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The Acanthodii, or acanthodians, were a group of jawed fish, usually considered a , and closely related to the . They are sometimes known as "spiny sharks", which is half right - they were spiny enough, but they were not sharks.

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Acanthodes

Range: Ordovician to Early Permian.

Phylogeny: : + *: Climatiiformes + (Ischnacanthiformes + Acanthodiformes).

Update: Hanke & Wilson (2006) have redescribed Brochoadmones, an acanthodian of uncertain affinities, on the basis of superlative material from the MOTH locality in Northerrn Canada. As originally reported, it looked a great deal like our image of Acanthodes. Among many new and interesting features, two points stand out. First, in addition to the ventral spine pairs, Brochoadmones has a thin, very osteichthyan pectoral fin, located about midway up the body wall. The fin has a short pectoral fin spine (much shorter than the fin) "resembling a bivalve shell" but no ossified endoskeleton. The implication seems to be that the osteichthyan pectoral fin is only indirectly related to the usual ventral fin spines of the acanthodians. Second, the head turns out to be very small, the fish being deep-bodied and almost trapezoidal overall. ATW061003.

Characters: Crania based on 4 large plates bridged by cartilage; subterminal mouth; branchiostegal rays present; no enamel on teeth, teeth not regularly replaced; jaws of dermal bone; orbits large and far anterior, with circumorbital plates; bony operculum; branchiostegal rays present; otoliths in inner ear; narrow-based braincase with a transverse ventral fissure; braincase & otic capsule well-ossified; primitively, several opercular scales; vertebral centra not ossified; no ribs; heterocercal tail; fin spines on all except caudal fins; generally membranous fins without rays except for spines; various articulations of pectoral spine with shoulder girdle; up to 6 paired intermediate spines between pectoral and pelvic fins; scales like , but no pulp cavity: base bulges & new odontodes of dentine or mesodentine surround and overgrow old with layered effect; fins covered with similar scales, like lepidotrichia of Osteichthyes; cranial large scales not homologous to Osteichthyes.

Links: ; ; ; ; ; (Slovenian: apparantly a nice summary); (Swedish: introduction to the Osteichthyes with a brief section on the Acanthodii); (the usual amazing images from this site); by Zaijicat Publications). APW 040806. HAJ100226

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Climatius

Range: to .

Phylogeny: Acanthodii: (Ischnacanthiformes + Acanthodiformes) + *.

Characters: May be paraphyletic basal group. Generally small-bodied (<15 cm); teeth minute & petalliform, not shed; extramandibular absent; palatoquadrate elongate & without otic articulation; 1 large and several small gill covers; broad-based spines with ridges; 2 dorsal fins; 2+ intermediate spines between pectoral and pelvic fins; well-developed dermal pectoral plates.

Links: (Italian); ; (Japanese). HAJ100226

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Ischnacanthus

Range: to

Phylogeny: Acanthodii:: Acanthodiformes + *.

Characters: Large toothed bones on palatoquadrate (looks like a shark); dermal pectoral covering reduced; 1 large gill cover; fin spines slender, with orthodentine superficial layer; some large (2m) species. APW 040806.

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Acanthodes, Howattacanthus

Range: to

Phylogeny: Acanthodii:: Ischnacanthiformes + *.

Characters: Extramandibular bone present; large orbits; well-developed gill rakers; 1 dorsal fin; 1 very large gill cover; no teeth; no intermediate spines between pectoral and pelvic fins; some elongated forms.

Links: ; ; . APW 040806, HAJ100226

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Acanthodii
   |--Climatiiformes
   `--+--Ischnacanthiformes
      `--Acanthodiformes

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  • Hanke, GF & MVH Wilson (2006), Anatomy of the Early Devonian acanthodian Brochoadmones milesi based on nearly complete body fossils, with comments on the evolution and development of paired fins. J. Vert. Paleontol. 26: 526-537. Acanthodii.
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