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Placoderm topics: | | | |


Acanthothoraci - Ancestral Placoderms?


Two order viagra placoderms, and of , from the () of South-East . Graphic © ‎Stanton F. Fink


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The Acanthothoraci ("spiny chests") were a group of -like who were closely related to the placoderms. Superficially, the acanthoracids resembled scaly chimaeras, lightly armored , or (relatively) heavily armored . They were distinguished from chimaeras by the presence of large scales and plates, tooth-like beak plates, and the typical bone-enhanced placoderm eyeball. They were distinguished from other placoderms due to differences in the anatomy of their skulls, and due to patterns on the skull plates and thoracic plates that are unique to each order.


(The following menu and phylogeny refers to subtopics of this page)


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Even more than with any other placoderm groups, are highly controversial.

One theory is that the acanthothoracids were closely related to the rhenanids because of similar arrangements of the plate armor, and that the patterns on the former's plates resemble the patterns of the latter's tubercle mosaics. Some authorities unite the two groups, though, they differ in which order to place the other in. According to 1980, 1986, the acanthothoracids can be considered primitive rhenanids on the basis of the dorsal (upper) position of their nostrils.

A very different hypothesis, by 1984, and followed by 1996 pp.170-2, holds that the Acanthothoraci, and not the , were the basal placoderms, and that the Acanthothoraci (as we currently know them) were . In this hypothesis, the acanthothoracids as a basal assemblage from which the other Placoderm lineages (the , the , and the ) emerged.

These two theories are actually not incompatible, since if the Acanthothoraci are a paraphytletic , there would still be those that evolve into - and hence represent primitive representatives and sister clades of - the Rhenanida.

The cladogram used here is based on that given by Janvier, following Goujet 1984, except that is here presented as a more basal Placoderm. This cladogram should not be taken as anything other than one possible interpretation among many.

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From what can be inferred from the mouthplates of fossil specimens, the acanthothoracids were ecologically similar to modern-day chimaeras, being a clique of shellfish hunters. Competition with their relatives, the ptyctodont placoderms, may have been one of the main reasons for the acanthothoracids' extinction prior to the Mid Devonian extinction event.

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?, Early to Mid

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  • Acanthothoraci , 1944 [Acanthothoraciformes]
    • Palaeacanthaspidae (author?) (year?)
      • Dobrowlania
      • Kimaspis
      • Kolymaspis
      • Kosoraspis
      • Palaeacanthaspis
      • Radotina [Holopetalichthys]
      • Romundina
    • Family Weejasperaspididae White, 1944
      • Murrindalaspis
      • Weejasperaspis

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  • Mikko's Phylogeny Archive

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  • (1984). Placoderm interrelationships: a new interpretation, with a short review of placoderm classification. Proceedings of the Linnean Society of New South Wales. 107, 211-241.
  • . Early Vertebrates Oxford, New York: Oxford University Press, 1996.
  • The Rise of Fishes: 500 Million Years of Evolution Baltimore: The John Hopkins University Press, 1996.
  • (1986). The relationships of placoderm fishes. Zoological Journal of the Linnean Society, 88, 1-57.

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