Annelida

From Palaeos

Protostomia
`--o Lophotrochozoa [or Spiralia]
   `--+--Bryozoa [Ectoprocta]
      |--Platyzoa
      `--o Trochozoa 
         |==Coeloscleritophora† ("Procoelomata†")
         |--Nemertea [Nemertinea]
         |--o Eutrochozoa
         |  |?-Tullimonsterida †
         |  |--Mollusca
         |  |--Hyolitha †
         |  |--Sipuncula 
         |  `--Annelida
         `--o Brachiozoa
            |--Phoronida 
            `--Brachiopoda

The Annelida or annelids (from Latin annellus - "little ring"), are a large phylum of animals, comprising the segmented worms, with about 15,000 modern species including the well-known earthworms and leeches. They are found in most wet environments, and include many terrestrial, freshwater, and especially marine species (such as the polychaetes), as well as some which are parasitic or mutualistic. They range in length from under a millimeter to over 3 meters (the seep tube worm Lamellibrachia luymesi).

Lophotrochozoa
`==Hallkeriida
   `--+--Wiwaxia corrugata †
      `--+--Canadia spinosa †
         `==Polychaeta (paraphyletic) (Annelida) 
            |--Myzostomida
            |--Pogonophora
            |--Vestimentifera
            `--+--+--Banffida †
               |  `--+-- Echiura [Echiuroidea] 
               `--Clitellata
                  |--Acanthobdellida
                  |--Branchiobdellida
                  |--Hirudinea
                  `--Oligochaeta       

[edit] Taxonomy

<==Annelida [Chaetopoda]
   |  i. s.: Aeolosomatida
   |         Acanthobdella peledina
   |         Dodecaceria concharum
   |         Neanthes
   |           |--N. arenaceodentata
   |           `--N. diversicolor
   |         Paranerilla limicola
   |--Pogonophora [Brachiata]
   |    |--Vestimentifera [Obturata]
   |    |    |--Riftia pachyptila
   |    |    `--Ridgeia piscesae
   |    |--Siboglinum [Perviata]
   |    |    `--S. fiordicum
   |    `--Sabellidites [Sabelliditida, Sabelliditidae]
   |         `--S. cambriensis
   |--Echiura [Echiuroidea]
   |    |  i. s.: Bonellia viridis
   |    |         Thalassema mellita
   |    |--Urechis [Xenopneusta]
   |    |    `--U. caupo Fischer & MacGintie 1928
   |    `--Echiuroinea
   |         |--Echiurus
   |         `--Ochetostoma erythrogrammon
   |--Clitellata
   |    |  i. s.: Calliobdella vivida
   |    |         Platybdella buccalis
   |    |         Piscicola
   |    |           |--P. funduli
   |    |           |--P. milneri
   |    |           `--P. punctata
   |    |--Oligochaeta
   |    `--Hirudinea
   `--Polychaeta [Pharyngata]
        |  i. s.: Hesionidae
        |         Sphaerodoridae
        |           |--Ephesia gracilis
        |           |--Sphaerodoropsis
        |           `--Sphaerodorum
        |                |--S. claparedei Greef 1866
        |                `--S. greefii
        |         Eumenia crassa
        |         Lumbrineris
        |           |--L. fragilis
        |           |--L. impatiens
        |           |--L. latrelli
        |           |--L. longifolia
        |           `--L. tenuis
        |         Pherusa
        |           |--P. affinis
        |           `--P. eruca
        |         Nephthys
        |           |--N. ciliata
        |           `--N. hombergii
        |         Ophelina
        |         Ninoe
        |         Dasybranchus caudatus
        |         Maldanidae
        |           |--Maldane sarsi
        |           |--Petaloproctus socialis
        |           |--Branchioasychis americana
        |           `--Clymenella
        |                |--C. torquata
        |                `--C. zonalis
        |         Scalibregma inflatum
        |         Amharete acutifrons
        |         Asabelides sibirica
        |         Pectinariidae
        |           |--Cistenides
        |           |    |--C. gouldii [=Pectinaria gouldii]
        |           |    `--C. hyperborea
        |           `--Pectinaria
        |                |--P. californiensis
        |                `--P. koreni
        |         Eulalia
        |           |--E. bilineata
        |           |--E. macroceros [=Pterocirrus macroceros]
        |           `--E. viridis (Linnaeus 1767)
        |         Glycinde armigera
        |         Laonice cirrata
        |         Magelona
        |           |--M. mirabilis
        |           `--M. sachalinensis
        |         Praxillella praetermissa
        |         Sigambra
        |           |--S. bassi
        |           `--S. hanaokai (Kitamori 1960)
        |         Spiophanes bombyx
        |         Tharyx pacifica
        |         Chone teres
        |         Pseudopotamilla occelata
        |         Jugaria quadriangularis
        |         Chrysopetalum occidentale
        |         Circeis
        |           |--C. armoricana
        |           `--C. spirillum
        |         Flabelligera affinis (see below for synonymy)
        |         Hermilepidonotus robustus
        |         Lepidonotus squamatus
        |         Eudistylia polimorpha
        |         Eumida
        |           |--E. sanquinea
        |           `--E. tubiformis
        |         Genetylli castanea
        |         Halosydna brevisetosa
        |         Odontosyllis maculata
        |         Pista
        |           |--P. cristata
        |           `--P. elongata
        |         Polymnia trigonostoma
        |         Demonax fullo
        |         Neoamphitrite grayi
        |         Nicolea gracilibranchis
        |         Alvinella pompejana
        |         Sabellaria
        |           |--S. alveolata
        |           |--S. cementarium
        |           `--S. spinulosa
        |         Euthalanessa dendrolepis
        |         Hermonia histrix
        |         Lagisca extenuata
        |         Polydontes maxillosus
        |         Mysta picta
        |         Sige limbata
        |         Kefersteinia cirrata
        |         Ophiodromus flexuosus (Delle Chiaje 1825) (see below for synonymy)
        |         Podarke pallida
        |         Syllidia armata
        |         Langheransia cornuta
        |         Trypanosyllis coeliaca
        |         Nematonereis unicornis
        |         Naineris laevigata
        |         Aonides oxycephala
        |         Aricidea
        |           |--A. cerrutii Laubier 1967 [=Acesta cerrutii]
        |           `--A. jeffreysii
        |         Phyllochaetopterus solitarius
        |         Raphidrilus nemasoma
        |         Ophelia
        |           |--O. limacina
        |           `--O. neglecta Schneider 1887
        |         Notomastus latericeus
        |         Owenia fusiformis
        |         Amphictene auricoma
        |         Ampharete grubei
        |         Melinna palmata
        |         Dialychone acustica
        |         Fabricia sabella
        |         Jasmineira candela
        |         Ditrupa arietina
        |         Praegeria remota
        |         Synelmis albini (Langerhans 1881)
        |         Pionosyllis augeneri Hartmann-Schröder 1979
        |         Branchiosyllis pacifica Rioja 1941
        |         Arenicolidae
        |           |--Arenicola
        |           `--Archarenicola
        |         Acholoe squamosa (Delle Chiaje 1825) (see below for synonymy)
        |         Adyte assimilis (McIntosh 1874) (see below for synonymy)
        |         Arctonoe vittata
        |         Antinoella sarsi [incl. Polynoe cirrata Grube 1876, Harmothoe cirrata]
        |         Hermadion hyalinus
        |         Subadyte pellucida (Ehlers 1864-1868) (see below for synonymy)
        |         Phylo foetida [=Aricia foetida]
        |         Amphitrite edwardsi
        |         Euclymene lombricoides
        |         Phalacrostemma cidariophilum
        |         Platynereis
        |           |--P. dumerilii
        |           `--P. megalops
        |         Burgessochaeta setigera
        |         Armandia brevis (Moore 1906)
        |         Hermodice carunculata
        |         Hesperonoe adventor
        |         Nicomache lumbricalis
        |         Pygospio
        |           |--P. elegans
        |           `--P. muscularis Ward 1981
        |         Sternaspis [Sternaspidae]
        |           `--S. scutata
        |--Archiannelida
        |--Spionida
        |--Capitellidae [Capitellida]
        |    |--Capitella capitata
        |    `--Heteromastus filiformis
        |--Eunicida
        |    |  i. s.: Eunice viridis
        |    |--Ramphoprionidae
        |    `--Dorvilleidae
        |--Sabellida
        |    |  i. s.: Sabella
        |    |           |--S. melanostigma
        |    |           `--S. pavonina
        |    |         Protula tubularia
        |    `--Serpulidae
        |--Psammodrilidae [Psammodrilida]
        |    |--Psammodriloides Swedmark 1961
        |    |    `--P. fauveli Swedmark 1961
        |    `--Psammodrilus Swedmark 1953
        |         |--P. aedificator Kristensen & Nørrevang 1982
        |         `--P. balanoglossoides Swedmark 1952
        |--Terebellida
        |    |  i. s.: Lanice conchilega
        |    `--Terebellidae
        |         |--Terebella ehrenbergi
        |         `--Polycirrus
        |              |--P. haematodes
        |              `--P. pallidus
        |--Orbiniida
        |    |--Questa Hartman 1966 [Questidae]
        |    |    `--Q. media Westheide 1981
        |    `--Orbiniidae
        |         |--Orbinia latreilli [=Aricia latreilli]
        |         |--Schroederella Laubier 1962 [Protoariciinae]
        |         |    |--S. berkeleyi Laubier 1971
        |         |    `--S. pauliani Laubier 1962
        |         `--Scoloplos armiger
        |--Cirratulida
        `--Phyllodocida
             |  i. s.: Aphrodita aculeata
             |--Pisionidae
             |--Phyllodocidae
             |--Syllidae
             |--Polynoidae
             |--Pholoe Johnston 1839 [Sigalionidae]
             |    `--P. swedmarki Laubier 1975
             |--Glycera [Glyceridae]
             |    |--G. alba
             |    |--G. americana
             |    |--G. capitata
             |    |--G. convoluta
             |    `--G. lapidum
             |--Goniadidae
             |    |--Progoniadides Hartmann-Schröder 1974
             |    |    `--P. laevis Hartmann-Schröder 1974
             |    |--Goniada
             |    |    |--G. maculata
             |    |    `--G. norvegica
             |    `--Goniadides Hartmann-Schröder 1960
             |         |--G. aciculata Hartmann-Schröder 1960
             |         `--G. falcigera Hartmann-Schröder 1962
             `--Nereididae

Acholoe squamosa (Delle Chiaje 1825) [incl. Polynoe astericola, A. astericola, P. asterinae, P. malleata Grube 1855]

Adyte assimilis (McIntosh 1874) [=Hermadion assimile, Scalisetosus assimilis; incl. H. echini Giard 1886]

Flabelligera affinis [=Siphonostoma affinis; incl. Chlorhema dujardinii Giard 1880, S. dujardini]

Ophiodromus flexuosus (Delle Chiaje 1825) [=Nereis flexuosa, Podarke flexuosa, Stephania flexuosa; incl. Ophiodromus vittatus Sars 1861]

Subadyte pellucida (Ehlers 1864-1868) [=Polynoe pellucida, Adyte pellucida, Hermadion pellucidum, Lepidonotus pellucidus, Scalisetosus pellucidus; incl. H. fragile Clarapède 1868, H. fugax Giard 1890, H. sabatieri Darboux 1900]

* Type species of genus indicated

[edit] References

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