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Taxonomy Very Speculative Phylogeny

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Anteosaurus


Anteosaurus magnificus dining on Illustration © Stephen Priestley


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Anteosaurus magnificus is a huge , known from the (Middle ) of south (now, South Africa). This giant lumbering predator, the size of a , was the largest carnivore and apex preditor of its environment. It no doubt preyed on the contemporary herbivorous . Very similar animals are known from eastern (China - ) and equatorial (Russia - , ) Pangea from more or less the same time. All these animals died out with the end-Capitanian turn-over.

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            • Genus: Anteosaurus Watson 1921
              • Species: magnificus Watson 1921 (

Locality:Beaufort West, Karoo deposits of South Africa

Age:

Material: 32 partial or complete skulls, assorted post-cranial elements

size: skull 50 to 80 cm long; overall length 3 to 5 meters

Possible distribution: Southern

Stratigraphic range: epoch

Lifestyle/Ecological role:

Size: adult skull - upto 80 cm long; overall length upto 5 m; weight upto 500-600 kg;

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Characters: postfrontal forms a boss of variable size overhanging the dorso-posterior border of the orbit [B69].

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Anteosaurus magnificus Watson 1921. Known from 32 partial or complete skulls, assorted post-cranial elements. Probably the only valid genus and species of Anteosaurus [B69].

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Note: On this basis of his 1969 description, Boonstra [B69] synonymised six of the seven genera named from the Tapinocephalus zone: Eccasaurus, Anteosaurus, Titanognathus, Dinosuchus, Micranteosaurus, and Pseudanteosaurus. Of these, he says, Dinosuchus and Titanognathus can safely be considered synonyms of Anteosaurus. Eccasaurus, with a holotype of which the cranial material consists of only few typical anteosaurid incisors, appears to be only determinable as to family. The skull fragment forming the holotype of Pseudanteosaurus can best be considered as an immature specimen of Anteosaurus. Micranteosaurus, the holotype of which contains a small snout, was previously considered a new genus on account of its small size but is better be interpreted as a young specimen of Anteosaurus. And likewise, the large number of species attributed to the genus Anteosaurus can also be considered synonyms.

Boonstra still considers as valid the genus Paranteosaurus, which is defined as a genus of anteosaurids in which the postfrontal is not developed to form a boss. This is probably an example of individual variation and hence another synonym of Anteosaurus.

As with the genera, so the species are also mostly synonyms. The oldest species name, Anteosaurus magnificus, is thus the only valid one. To quote Boonstra again:

"We have 32 skulls of Anteosaurus, of which 16 are reasonably well preserved and on them ten species have been named. To differentiate between the species the following main characters have been used: the number, size and shape of the teeth, skull size, shape and the nature of the pachyostosis. On re-examination it has become clear that the crowns of the teeth are seldom well preserved; basing the count for the dental formula on the preserved roots is unreliable. As this is affected by age and tooth generation; size of skull is a function of age and also possibly sex; skull shape is greatly affected by post-mortem deformation, and the variability in the pachyostosis, which may be specific in some respects, can just as well be the result of...physiological processes. Specific diagnosis consisting of the enumeration of differences of degree in features such as the above can hardly be considered as sufficient indication of the existence of discrete species.... A. magnificus thus has the following synonyms: abeli, acutirostrus, crassifrons, cruentus, laticeps, levops, lotzi, major, minor, minusculus, parvus, priscus and vorsteri."

The reasons for the synonymy he gives as follows: the holotypes of cruentus, levops, minor, minusculus, and parvus are of immature animals; lotzi and priscus are specifically indeterminable; and in the case of abeli, acutirostrus, crassifrons, laticeps, major and vorsteri because the characters used are either due to post-mortem deformation or individual variation in the degree of pachyostosis. Ironically, a lot of these specific and generic synonyms were named by Boonstra himself, who reversed his earlier decisions later on.

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The following list of synonyms is from Tracy Ford's Paleofiles -

Genus: Anteosaurus WATSON, 1921

  • Broomosuchus CAMP, TAYLOR & WELLES, 1942
  • Dinosuchus BROOM, 1936
  • Micranteosaurus BOONSTRA, 1954
  • Pseudanteosaurus BOONSTRA, 1954
  • Titanognathus BROILI & SCHOREDER, 1935

A. magnificus WATSON, 1921 (Type)

  • Titanosuchus ferox BROOM, 1910 non OWEN, 1876
  • Pseudanteosaurus minor (BROOM, 1929)
  • Anteosaurus minor BROOM, 1929
  • Anteosaurus lotzi (BROILI & SCHROEDER, 1935)
  • Titanognathus lotzi BROILI & SCHROEDER, 1935
  • Anteosaurus minusculus BOONSTRA, 1936
  • Anteosaurus minor BOONSTRA, 1936
  • Anteosaurus vorsteri (BROOM, 1936)
  • Dinosuchus vorsteri BROOM, 1936
  • Broomosuchus vorsteri (BROOM, 1936) CAMP, TAYLOR & WELLES, 1942
  • Anteosaurus abeli BOONSTRA, 1952
  • Anteosaurus major BOONSTRA, 1954
  • Micranteosaurus parvus BOONSTRA, 1954
  • Anteosaurus acutirostris BOONSTRA, 1954
  • Anteosaurus crassifrons BOONSTRA, 1954
  • Anteosaurus laticeps BOONSTRA, 1954
  • Anteosaurus cruentus BOONSTRA, 1954
  • Anteosaurus levops BOONSTRA, 1954
  • A. sp.

The following also probably belong to this species:

Genus: Eccasaurus BROOM, 1909

  • E. priscus BROOM, 1909 (Type)

Genus: Paranteosaurus BOONSTRA, 1954

  • P. primus BOONSTRA, 1954 (Type)

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(excellent images) (notes on way of life) (fauna) (abstract on fauna & biozone).

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Boonstra (1969) [B69]; Müller (1985) [M85].

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MAK000902, ATW030807.

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