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Arthropoda (see below for synonymy)
   |  i. s.: Dictyocaris Salter 1860
   |         Urokodia Hou, Chen & Lu 1989
   |         Tontoia Walcott 1912
   |         Retifacies Hou, Chen & Lu 1989
   |         Rhombicalvaria Hou 1987
   |         Helmetia Walcott 1917
   |         Corcorania Jell 1980
   |         Serracaris Briggs 1978
   |           `--S. lineata [=Anomalocaris lineata]
   |         Cambropodus gracilis
   |         Protocaris Walcott 1884
   |         Achanarraspis Anderson et al. 2000
   |         Arthrogyrinus Wilson & Almond 2001
   |           `--*A. platyurus Wilson & Almond 2001
   |--Sidneyia Walcott 1911 [Limulavida, Sidneyidea]
   |    `--S. inexpectans
      `--+--Yohoiidea [Protochelicerata, Yohoiidacea]
         |    |--Jiangfengia Hou 1987 [Jianfengiida]
         |    `--Yohoia Walcott 1912 [Yohoiida]
         |         `--Y. tenuis
         `--+--Sanctacaris Briggs & Collins 1988 [Sanctacarida, Sanctacaridea]
            |    |--‘Utahcaris’ orion Conway Morris & Robison 1988
            |    `--Sanctacaris uncata Briggs & Collins 1988
               |  `--+--Occacaris
               |     `--+--Clypecaris
               |        `--+--+--Pectocaris
               |           |  `--Fuxianhuiida [Fuxianhuiata]
               |           |       |--Fuxianhuia protensa Hou 1987
               |           |       `--Changjiangocaris longiformis Hou & Bergström 1991
               |           `--Canadaspidida
               |                |  i. s.: Rehbachiella kinnekullensis Müller 1983
               |                |--Perspicaris Briggs 1977
               |                `--+--Canadaspis Novoshilov 1960
               |                   |    `--C. perfecta (Walcott 1912)
               |                   `--Brachiocaridea
               |                        |--Odaraia Walcott 1912 [Odaraiata, Odaraiida]
               |                        |    `--O. alata
               |                        `--Branchiocaris Briggs 1976 (see below for synonymy)
               |                             `--B. pretiosa (Resser 1929) [=Protocaris pretiosa]
               `--Euarthropoda [Emeraldellacea]
                    `--+--Emeraldella Walcott 1912 [Emeraldellida]
                          `--Mandibulata [Antennata, Atelocerata, Biantennata, Tracheata]
                               |  i. s.: Euthycarcinoidea [Archicopepoda, Euthycarcinata]
                               |           |  i. s.: Synaustrus brookvalensis (Riek 1964)
                               |           |         Schramixerxes gerem (Schram & Rolfe 1982)
                               |           |         Kalbarria brimmellae
                               |           |--Sottixeriformes
                               |           |    |--Pieckoxerxes piekoae (Schram & Rolfe 1982)
                               |           |    `--Sottyxerxes multiplex Schram & Rolfe 1982
                               |           `--Euthycarciniformes
                               |                |  i. s.: Euthycarcinus
                               |                |           |--E. ibbenburensis
                               |                |           `--E. kessleri Handlirsch 1914
                               |                `--Kottixerxidae
                               |                     |--Kottixerxes Schram 1971
                               |                     |    |--*K. gloriosus Schram 1971
                               |                     |    `--K. anglicus Wilson & Almond 2001
                               |                     `--Smithixerxes Schram & Rolfe 1982
                               |                          |--*S. juliarum Schram & Rolfe 1982
                               |                          `--S. pustulosus Wilson & Almond 2001
                               |         Marrellomorpha
                               |           |--Vachonisia Lehmann 1955 (see below for synonymy)
                               |           `--Marrellidea
                               |                |--Mimetaster Gurich 1931 (see below for synonymy)
                               |                `--Marrella Walcott 1912 [Marrellata, Marrellida]
                               |                     `--M. splendens
                               `--Pancrustacea [Tetraconata]

Arthropoda [Burgessidea, Canadaspididea, Cheliceromorpha, Emeraldellidea, Euthycarcinomorpha, Merostomoidea, Schizoramia, Trilobitoidea, Trilobitomorpha]

Branchiocaris Briggs 1976 [Branchiocarata, Branchiocarida]

Mimetaster Gurich 1931 [Mimetasterata, Mimetasterida]

Vachonisia Lehmann 1955 [Vachonisiata, Vachonisiida]

* Type species of genus indicated

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Bousfield, E. L. 1995. A contribution to the natural classification of Lower and Middle Cambrian arthropods: Food-gathering and feeding mechanisms. Amphipacifica 2: 3-34.

Briggs, D. E. G., B. S. Lieberman, S. L. Halgedahl & R. D. Jarrard. 2005. A new metazoan from the Middle Cambrian of Utah and the nature of the Vetulicolia. Palaeontology 48 (4): 681-686.

Budd, G. E. 2002. A palaeontological solution to the arthropod head problem. Nature 417: 271-275.

Conway Morris, S. 1998. The Crucible of Creation. Oxford University Press: Oxford.

Giribet, G., G. D. Edgecombe & W. C. Wheeler. 2001. Arthropod phylogeny based on eight molecular loci and morphology. Nature 413: 157-161.

Gould, S. J. 1989. Wonderful Life. Vintage: London.

Lange, S., & F. R. Schram. 2002. Possible lattice organs in Cretaceous Thylacocephala. Contributions to Zoology 71 (4): 159-169.

Newman, M. J. 2002. A new naked jawless vertebrate from the Middle Devonian of Scotland. Palaeontology 45 (5): 933-941.

Rode, A. L., & B. S. Lieberman. 2002. Phylogenetic and biogeographic analysis of Devonian phyllocarid crustaceans. Journal of Paleontology 76 (2): 271-286.

Waloszek, D., & J. A. Dunlop. 2002. A larval sea spider (Arthropoda: Pycnogonida) from the Upper Cambrian ‘Orsten’ of Sweden, and the phylogenetic position of pycnogonids. Palaeontology 45 (3): 421-446.

Whittington, H. B., & D. E. G. Briggs. 1985. The largest Cambrian animal, Anomalocaris, Burgess Shale, British Columbia. Philosophical Transactions of the Royal Society of London 309: 569-609.

Wilson, H. M., & J. E. Almond. 2001. New euthycarcinoids and an enigmatic arthropod from the British coal measures. Palaeontology 44 (1): 143-156.



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