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Coccosteus is an extinct genus of . Its fossils have been found throughout Europe and the US. The majority of these have been found in freshwater sediments, though, such a large range suggests that they may have been able to enter saltwater. The largest specimens were about 40 centimers, though, most tended to be about 20 to 24 centimeters in length.

Like all other , Coccosteus had a joint between the armor of the body and . In addition to that, it also had an internal joint between its neck and the back of the skull, allowing for the mouth to be opened even wider. Along with the longer jaws, this allowed Coccosteus to feed on fairly large prey. The up-and-down movement of the skull also allowed for more water to be pumped through the . Possibly, the creature supplemented its diet with organic material filtered from mud using the gills. As with all other arthrodires, Coccosteus had bony dental plates embedded in its jaws, forming a beak. The beak was kept sharp by having the edges of the dental plates grind away at each other. Overall the creature looked a lot like its gigantic cousin , save that its eyes were closer to the end of its snout than its larger relative.

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