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Coleoidea comprise a subclass of in which the shell is absent or internal and which have upwards of ten arms, in which case two are termed tentacles, in as many as five pairs which protrude from the head and encircle the mouth. Recent Coleoids are dibranchiate, that is have two gills as opposed to the four found in recent represented by Nautilus. Coleoids are also referred to as neocephalopods.

The Coleoidea has been divided into two principle groups, infraclasses or superorders, the Belemnoidea and Neocoleoidea. Recent coleoids are Neocoloidea and are further divided into the ten armed Decapodiformes which includes squids, octopods, cuttle-fishes, and Spirula, and the eight armed Octopodiformes, octopuses.

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Subclass Coleoidea

, the genus Spirula

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   |  i. s.: Onychoteuthis borealijaponicus AFL84
   |         Gonatus [Gonatidae] W81
   |           `--G. fabricii W81
   |         Illex W81
   |           |--I. coindeti B26
   |           `--I. illecebrosus W81 [=Ommastrephes illecebrosus B26]
   |         Dosidicus gigas W81
   |         Phragmoteuthidida S64
   |         Bactritimimus P68
   |           |--B. girtyi Flower & Gordon 1959 P68
   |           `--B. ulrichi Flower & Gordon 1959 P68
   |         Eobelemnites caneyense Flower 1945 P68
   |         Hematites P68
   |           |--H. barbarae Flower & Gordon 1959 P68
   |           `--H. burbankensis Flower & Gordon 1959 P68
   |         Paleoconus bakeri Flower & Gordon 1959 P68
   |--Belemnitida [Belemnoidea] P98
   |    |--Zugmontites mojsisovicsi Reis 1907 E64
   |    `--Atractites E64
   |         |--A. breviconus Reis 1907 E64
   |         |--A. conicus Mojsisovics 1871 E64
   |         |--A. convergens Hauer 1847 E64
   |         |--A. ellipticus Mojsisovics 1871 E64
   |         |--A. giganteus Gemmellaro 1904 E64
   |         `--A. meneghinii Salomon 1895 E64
   `--+-- GO06
      `--+--Stauroteuthis syrtensis GO06
         `--+--Vampyroteuthis infernalis Chun 1903 GO06, RP07
            `--+--Sepia [Sepiida, Sepiidae, Sepioidea] GO06
               |    |--S. apama Gray 1849 F27
               |    |--S. cultrata Hoyle 1885 H09
               |    |--S. elegans GW02
               |    |--S. esculenta Hoyle 1886 H09
               |    |--S. latimanus Quoy & Gaimard 1832 BW09
               |    |--S. officinalis GO06
               |    |--S. pfefferi Hoyle 1885 H09
               |    `--S. pharaonis Ehrenberg 1831 BW09
               `--+--+--Heteroteuthis hawaiiensis GO06
                  |  `--+--Rossia palpebrosa GO06
                  |     `--Sepiola GO06
                  |          |--S. affinis GO06
                  |          |--S. atlantica NN03
                  |          |--S. intermedia NN03
                  |          |--S. ligulata NN03
                  |          |--S. robusta NN03
                  |          `--S. rondeleti NN03
                  `--+--Cranchia scabra GO06
                     `--+--+--Idiosepius pygmaeus GO06
                        |  `--Loliginidae BW09
                        |       |--Loligo GO06
                        |       |    |--L. bleekeri NS03
                        |       |    |--L. opalescens W81
                        |       |    |--L. pleii Blainville 1823 HC79
                        |       |    `--L. vulgaris H87
                        |       |--Sepioteuthis lessoniana Férussac 1831 BW09
                        |       `--Photololigo BW09
                        `--+--+--Histioteuthis GO06
                           |  |    |--H. bonellii B70
                           |  |    `--H. hoylei GO06
                           |  `--+--Architeuthis dux GO06
                           |     `--Lepidoteuthis grimaldii GO06
                           `--+--Spirula [Spirulidae] GO06
                              |    `--*S. spirula (Linnaeus 1758) GO06, H09 [=Nautilus spirula H09]
                              `--+--Moroteuthis GO06
                                 |    |--M. knipovitchi GO06
                                 |    `--M. robusta W81
                                 `--Ommastrephes GO06
                                      |--O. bartrami GO06
                                      |--O. oualaniensis Lesson 1830 H09
                                      `--O. sloani W81

* Type species of generic name indicated

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[AFL84] Antonelis, G. A., Jr., C. H. Fiscus & R. L. de Long. 1984. Spring and summer prey of California sea lions, Zalophus californianus, at San Miguel Island, California, 1978-79. Fishery Bulletin 82 (1): 67-76.

[B26] Bigelow, H. B. 1926. Plankton of the offshore waters of the Gulf of Maine. Bulletin of the Bureau of Fisheries 40 (2): 1-509.

[B70] Briggs, J. C. 1970. A faunal history of the North Atlantic Ocean. Systematic Zoology 19 (1): 19-34.

[BW09] Bryce, C. & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177-208.

[E64] Erben, H. K. 1964. Bactritoidea. In Treatise on Invertebrate Paleontology pt. K. Mollusca 3. Cephalopoda – General Features – Endoceratoidea – Actinoceratoidea – Nautiloidea – Bactritoidea (R. C. Moore, ed.) pp. K491-K505. The Geological Society of America and the University of Kansas Press.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320-485.

[GO06] Giribet, G., A. Okusu, A. R. Lindgren, S. W. Huff, M. Schrödl & M. K. Nishiguchi. 2006. Evidence for a clade composed of molluscs with serially repeated structures: Monoplacophorans are related to chitons. Proceedings of the National Academy of Sciences of the USA 103 (20): 7723-7728.

[GW02] Giribet, G., & W. Wheeler. 2002. On bivalve phylogeny: A high-level analysis of the Bivalvia (Mollusca) based on combined morphology and DNA sequence data. Invertebrate Biology 121 (4): 271-324.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H87] Holland, C. H. 1987. The nautiloid cephalopods: a strange success story. Journal of the Geological Society 144 (1): 1-15.

[HC79] Huff, J. A., & S. P. Cobb. 1979. Penaeoid and sergestoid shrimps (Crustacea: Decapoda). Memoirs of the Hourglass Cruises 5 (4): 1-101.

[NS03] Nardi, F., G. Spinsanti, J. L. Boore, A. Carapelli, R. Dallai & F. Frati. 2003. Hexapod origins: Monophyletic or paraphyletic? Science 299: 1887-1889.

[NN03] Nishiguchi, M. K., & V. S. Nair. 2003. Evolution of symbiosis in the Vibrionaceae: A combined approach using molecules and physiology. International Journal of Systematic and Evolutionary Microbiology 53: 2019-2026.

[P98] Prothero, D. R. 1998. Bringing Fossils to Life: An introduction to paleobiology. WCB McGraw-Hill: Boston.

[P68] Purnell, L. R. 1968. Catalog of the Type Specimens of Invertebrate Fossils. Part I: Paleozoic Cephalopoda. United States National Museum Bulletin 262: 1-198.

[RP07] Rousset, V., F. Pleijel, G. W. Rouse, C. Erséus & M. E. Siddall. 2007. A molecular phylogeny of annelids. Cladistics 23: 41-63.

[S64] Sweet, W. C. 1964. Cephalopoda - general features. In Treatise on Invertebrate Paleontology pt. K. Mollusca 3. Cephalopoda – General Features – Endoceratoidea – Actinoceratoidea – Nautiloidea – Bactritoidea (R. C. Moore, ed.) pp. K4-K13. The Geological Society of America and the University of Kansas Press.

[W81] Watson, L. 1981. Sea Guide to Whales of the World. Hutchinson: London.


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