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Ctenocystoidea are a small class of from the of western North America and Bohemia, first described in 1969. Dominguez-Alonso (1999); Parsley (1999). An undescribed form from the of Wales has also been assigned to the Ctenocystoidea, but preliminary illustrations (Dominguz-Alonso, 1999) are significantly different from other members of the class, so we'll have to wait for a detailed description before passing judgment.

Ctenocystoidea differ from the more basal in lacking an appendage. They are distinctly flattened and rectangular to trapezoidal in dorsal view. Like , they have a central covering of small plates with a row of much larger marginal plates around the outside edge of the .

The most distinctive feature of the Ctenocystoidea (and the source of their name) is the along the anterior margin. This is composed of two series (one each side of the mouth) of ctenoid ("comb-like") plates separated by paired plates on the anterior mid-line. The ctenoid plates were dorsally articulated and so capable of slight movement. A space behind the ctenoid organ held the . Dominguez-Alonso (1999) suggested that the ctenoid plates were covered in life by ciliary so the ctenoid organ acted as a drawing water into the ambulacrum.

The is located at the anterior right of the dorsal , while the pyramidal anus is located at the centre of the posterior margin. The mechanism by which ctenocystoids moved is problematical. Parsley (1999) suggested that the ctenoid plates may have been mobile enough to claw the animal forward. Rather more dramatically, it has been suggested that water may have been forcibly expelled through the anus, moving the animal by jet propulsion. Dominguez-Alonso (1999). The potential here for comedic commentary being obvious, we leave it to the reader's own interpretation.

Placing the Cincta and Ctenocystoidea in the phylogeny requires us to identify how the bilateral ambulacra of these classes correspond to the pentaradial ambulacra of crown echinoderms. Echinoderm ambulacra are identified by letters A-E, with A the anterior anbulacrum and the letters rising clockwise. In crown echinoderms, the ambulacra all run direct to the mouth, but in some stem echinoderms such as the , the B and C ambulacra fuse a short distance distally from the mouth, as do the D and E ammbulacra, forming the 2-1-2 ambulacral pattern. Jefferies (1997). In Cincta and Ctenocystoidea, the left is therefore probably the DE ambulacrum while the right is the BC ambulacrum. Dominguez-Alonso (1999) placed the Ctenocystoidea crownward of the the Cincta due to the loss of the posterior appendage and the greater symmetry of the ambulacra. We're willing to run with this hypothesis for now, but would like to point out that these are characters that could have easily appeared convergently.

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