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The basic framework of this tree in regard to the living taxa - (, (, (Hemichordata, Echinodermata))) - is probably fairly well-accepted at present. The placement of the fossil taxa is a little more contentious. The , shown here as stem-group deuterostomes, may be closer to chordates, or may not be deuterostomes at all (Briggs, Lieberman et al., 2005). The have been suggested to be either closer to chordates (even stem-group ), or stem-group hemichordates. Note also that this tree shows a definite minority view (endorsed, for instance, by Cavalier-Smith, 1998) that excludes from Chordata sensu stricto and restricts the latter name to the + clade.

Deuterostomia [Dexiothetica]
  |--+--+--Xenoturbella Westblad 1949 [Xenoturbellida]
  |  |  |    `--*X. bocki Westblad 1949
  |  |  `--Ambulacraria [Coelomopora]
  |  |       |--
  |  |       `--
  |  `--+--Nectocaris pteryx Conway Morris 1976
  |     `--+--Metaspriggina
  |        |--Pikaia [Pikaiidae]
  |        |    `--P. gracilens Walcott 1911
  |        `--Chordonia
  |             |--
  |             `--
       |  i. s.: Yuyuanozoon
       |--Yunnanozoa [Yunnanozoida, Yunnanozoonidae]
       |    |--*Yunnanozoon lividum Hou et al. 1991
       |    `--Haikouella Chen, Huang & Li 1999
       |         |--*H. lanceolata
       |         `--H. jianshanensis Shu, Conway Morris et al. 2003
            |--Pomatrum ventralis
            |--Xidazoon Shu, Conway Morris & Zhang in Shu, Conway Morris et al. 1999
            |    `--*X. stephanus Shu, Conway Morris & Zhang in Shu, Conway Morris et al. 1999
            |--Didazoon Shu & Han in Shu, Conway Morris et al. 2001
            |    `--*D. haoae Shu & Han in Shu, Conway Morris et al. 2001
            |--Skeemella Briggs, Lieberman et al. 2005
            |    `--*S. clavula Briggs, Lieberman et al. 2005
            |    |--V. cuneata Hou 1987 [=V. cuneatus]
            |    `--V. rectangulata
                 |--B. confusa Chen & Zhou 1997
                 `--B. constricta

* Type species of genus indicated

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Bourlat, S. J., C. Nielsen, A. E. Lockyer, D. T. J. Littlewood & M. J. Telford. 2003. Xenoturbella is a deuterostome that eats molluscs. Nature 424: 925-928.

Briggs, D. E. G., B. S. Lieberman, S. L. Halgedahl & R. D. Jarrard. 2005. A new metazoan from the Middle Cambrian of Utah and the nature of the Vetulicolia. Palaeontology 48 (4): 681-686.

Cavalier-Smith, T. 1998. A revised six-kingdom system of life. Biological Reviews 73: 203-266.

Dzik, J. 1995. Yunnanozoon and the ancestry of chordates. Acta Palaeontologica Polonica 40: 341-360.

Gould, S. J. 1989. Wonderful Life. Vintage: London.

Shu, D., S. Conway Morris, X.-L. Zhang, L. Chen, Y. Li & J. Han. 1999. A pipiscid-like fossil from the Lower Cambrian of south China. Nature 400: 746-749.

Shu, D.-G., S. Conway Morris, J. Han, L. Chen, X.-L. Zhang, Z.-F. Zhang, H.-Q. Liu, Y. Li & J.-N. Liu. 2001. Primitive deuterostomes from the Chengjiang Lagerst├Ątte (Lower Cambrian, China). Nature 414: 419-424.

Shu, D., S. Conway Morris, Z. F. Zhang, J. N. Liu, J. Han, L. Chen, X. L. Zhang, K. Yasui & Y. Li. 2003. A new species of yunnanozoan with implications for deuterostome evolution. Science 299: 1380-1384.



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