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Drosophilidae is one of two "fruit" fly families, though, technically, they're also known as "vinegar" or "pomace" flies. Unlike the "true" fruit flies, of , most drosophilid fruit flies lay their eggs in rotting fruit, or fungal matter, with a few Hawaiian species parasitizing other insects.

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Drosophilidae
   |--Scaptomyza fuscitarsus
   |--Paraleucophenga Hendel 1914
   |    |--*P. invicta (Walker 1857) [=Leucophenga invicta]
   |    |--P. emeiensis (Sidorenko 1998)
   |    |--P. javana (Okada 1988)
   |    |--P. neojavanai Singh & Negi 1992
   |    |--P. semiplunata (Duda 1939)
   |    |--P. shanyinensis (Chen & Toda 1994)
   |    |--P. shimai (Okada 1988)
   |    `--P. todai Fartyal & Singh 2004
   |--Scaptodrosophila Duda 1923
   |    |--*S. scaptomyzoidea Duda 1923
   |    `--S. latifasciaeformis
   `--Drosophila Fallén 1823 (see below for synonymy)
        |  i. s.: D. affinis
        |         D. aldrichi
        |         D. algonquin
        |         D. americana
        |           |--D. a. americana
        |           `--D. a. texana
        |         D. ananassae
        |         D. artificialis
        |         D. athabasca
        |         D. azteca
        |         D. capricorni
        |         D. earlei
        |         D. equinoxialis
        |         D. fumipennis
        |         D. guttifera
        |         D. heteroneura
        |         *Tanygastrella’ hypopygialis Duda 1924
        |         D. lacicola
        |         D. macrospina
        |           |--D. m. macrospina
        |           |--D. m. limpiensis
        |           `--D. m. ohioensis
        |         D. mauritiana
        |         *Adrosophila’ minuta Séguy 1938
        |         D. miranda
        |         D. montana
        |         D. mulleri
        |         *Spuriostyloptera’ multipunctata Duda 1923
        |         D. nasutoides
        |         D. nebulosa
        |         D. neorepleta
        |         D. nigrospiracula
        |         D. novamericana
        |         D. orientacea
        |         D. paulistorum
        |         D. persimilis
        |         D. prosaltans
        |         D. quadrilineata de Meijere 1911 [=*Chaetodrosophilella quadrilineata]
        |         D. saltans
        |         D. sechellia
        |         D. silvestris
        |         D. sturtevanti
        |         D. subobscura
        |         D. sulcata
        |         D. teissieri
        |         D. transversa
        |         D. tropicalis
        |         D. vibrissina
        |         D. willistoni
        |         D. (Paradrosophila Duda 1923)
        |           `--D. (*P.) pictipennis Kertész 1901
        |         D. (Spinulophila Duda 1923)
        |           `--D. (*S.) signata (Duda 1923) [=*Spinulophila signata]
        |--D. (Dorsilopha) busckii
        `--+--
           `--D. (Sophophora)
                |--+--D. (S.) obscura
                |  `--D. (S.) pseudoobscura
                `--+--D. (S.) erecta
                   |--D. (S.) melanogaster Meigen 1830
                   |--D. (S.) simulans
                   `--D. (S.) yakuba

Drosophila Fallén 1823 [incl. Adrosophila Séguy 1938, Chaetodrosophilella Duda 1923, Chaetodrosophila (l. c.), Spuriostyloptera Duda 1923, Tanygastrella Duda 1924]

* Type species of genus indicated

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Campbell, N. A. 1996. Biology (4th ed.) The Benjamin/Cummings Publishing Company, Inc.: Menlo Park (California).

Dayrat, B., C. Schander & K. A. Angielczyk. 2004. Suggestions for a new species nomenclature. Taxon 53 (2): 485-491.

Dobzhansky, T. 1937. Genetics and the Origin of Species. Columbia University Press: New York.

Dobzhansky, T. 1951. Genetics and the Origin of Species, 3rd ed. Columbia University Press: New York.

Fartyal, R. S., & B. K. Singh. 2004. A new species of the genus Paraleucophenga from the Kumaon region, India (Insecta, Diptera, Drosophilidae). Senckenbergiana Biologica 83 (2): 177-180.

Giordano, R., J. J. Jackson & H. M. Robertson. 1997. The role of Wolbachia bacteria in reproductive incompatibilities and hybrid zones of Diabrotica beetles and Gryllus crickets. Proceedings of the National Academy of Sciences of the USA 94: 11439-11444.

Gorlov, I. P., & N. Tsurusaki. 2000. Analysis of the phenotypic effects of B chromosomes in a natural population of Metagagrella tenuipes (Arachnida: Opiliones). Heredity 84: 209-217.

Hwang, U. W., M. Friedrich, D. Tautz, C. J. Park & W. Kim. 2001. Mitochondrial protein phylogeny joins myriapods with chelicerates. Nature 413: 154-157.

Kreitman, M., & H. Akashi. 1995. Molecular evidence for natural selection. Annual Review of Ecology and Systematics 26: 403-422.

Li, W.-H., & D. Graur. 1991. Fundamentals of Molecular Evolution. Sinauer: Sunderland (MA).

Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: An annotated species list. DSIR Entomology Division Report 3: 1-98.

Mayr, E. 1947. Ecological factors in speciation. Evolution 1: 263-288.

Noor, M. A. F. 2003. Genes to make new species. Nature 423: 699-700.

Robe, L. J., V. L. S. Valente, M. Budnik & E. L. S. Loreto. 2005. Molecular phylogeny of the subgenus Drosophila (Diptera, Drosophilidae) with an emphasis on Neotropical species and groups: a nuclear versus mitochondrial gene approach. Molecular Phylogenetics and Evolution 36: 623-640.

Singh, B. K., S. Dash & R. S. Fartyal. 2004. Revision of the species of the subgenus Drosophila (Drosophila) of the Kumaon region, India, with the description of eight new species. Senckenbergiana Biologica 83 (2): 163-176.

Slatkin, M. 1985. Gene flow in natural populations. Annual Review of Ecology and Systematics 16: 393-430.

Stebbins, G. L, & F. J. Ayala. 1981. Is a new evolutionary synthesis necessary? Science 213: 967-971.

Vogler, A. P., & R. Desalle. 1994. Diagnosing units of conservation management. Conservation Biology 8 (2): 354-363.

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