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The Ellesmerocerida are characterized by small to tiny straight and curved shells with very close spaced septa and a relatively large, staight ventral siphuncle with thick, typically concave, conntecting rings and generally short straight septal necks, in which cross positioned diaphragms are common. Concave rings swell inwardly so as to leave indentations, concavities, in internal siphuncle molds.

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The Ellesmerocerida comprise an order of Nautiloid Cephalopods included in the superorder Plectronoceratoidea and are derived from the Plectronocerida in the Late Cambrian. The Ellesmerocerida are divided into two suborders, the Ellesmeroceratina and the .


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The Ellesmerocerida have their origin in the stage of the Late , and are represented from that time by some 18 or so genera in four families found in the Fengshan Formation in China, of which three are limited to the Cambrian. First to appear are the Ellesmeroceratidae, starting with the early Trempealeauan Hunyuanoceras from the upper Yenchou Member. Hunyuanoceras has not been recorded above but the Ellesmeroceratidae diversify into some 12 genera in the overlying mid to late Trempealeauan Wanwankou Member. But only two, and are known to have survived the extenction before the end of the Cambrian and to have persisted into the Ordovician.

The ellesmeroceratids gave rise to two smaller families in the early Trempealeauan, the Acaroceratidae and Huaiheceratidae. The Acaroceratidae, containng two genera, are characterized by thin connecting rings and few diaphrams, and should perhaps be removed from the order. The Huaiheceratidae with three known genera and compressed exogastric forms. Both families are found throughout the Wanwankou. The fourth family, the Xiashanoceratidae, soley represented by the depressed exogastric Xiashanoceras (broader than high) found only in the Wanwankou.

No genus from the three minor families is known above the early late Trampealeauan and non is recorded from the late late Trampealeauan Chiushuko Member at the top of the formation that underlies the Ordovician.

Late Cambrian Ellesmeroceratidae are mostly orthoconic, although Chabactinoceras and Pseudendoceras are slightly endogastric and Eburoceras is extreemly so; unless Eburoceras is somehow exogastric with a dorsal siphuncle. A few are long and propotionally narrow as with the straight Tanycameroceras and the arcuately curved Eburoceras

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The Ellesmeroceratidae (Ellesmerocerida) recovered in the Gasconadian and greatly expanded and diversified. The Gasconadian is the lowest or earliest stage of the (Lower) Ordovcian, known especially in North America, as the Canadian.

From the narow orthoconic Ectenolites and the compressed and slightly endogastric Clarkoceras different forms evolved, combining straight to strongly curved and slender to rather stout aspects. Staight forms vary from the slender Ectonolites to the rapidly expanding Conocerina with Ellesmeroceras and Eremoceras in between. Curved forms vary from the slender and slightly curved Stemptonoceras though the moderately stout and endogastric Dakeoceras to the stout and compressed, strongly endogastric Levisoceras.

Later in the Gasconadian additional modifications showed up, mainly in the aperture which in some, such as the endgastirc Barnseoceras and Buehleroceras became constricted. Other developments include the broad grooves that run along the flanks of Oneotoceras.

Also from the Gasconadian are the ellesmeroceratid Pachendocera which gave rise to the through Proendoceras and Bassleroceras which belongs in the Bassleroceratidae and which gave rise to the

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The Ellesmeroceratidae continued to the end of the Lower Ordovician (Canadian), although as a minor contribution to the total cephalopod fauna. Also continuing to the end of the Canadian are the Bassleroceratidae and Protocycloceratidae which have their beginnings in the Gasconadian ellesmeroceratids. The Protocycloceratidae apparently gave rise to the Cyclostomiceratidae in the Late Canadian (Cassinian). All belong to the Ellesmeroceratina, except perhaps the Bassleroceratidae which Rousseau Flower included in the Tarphycerida.

Also, with its beginning early in the Middle Canadian, is the Eothinoceratidae, first of the Cyrtocerinina. Eothinoceras, the sole known genus of the family gave rise in the Late Canadian to Bathmoceras the sole genus of the Bathmoceratidae which stands as a possible origin of the ., and to the Late Ordovician

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The Ellesmerocerida are the origin of three and perhaps four orders, the Endocerida and Tarphycerida early in the Middle Canadian and the Orthocerida (Michelinocerida Flower) in the Late Canadian (Cassinian), unless the Baltoceratidae are included in which case that origin drops back into the Middle Canadian. The derivation of the Actinocerida remains uncertain although it most likely lies in the Ellesmerocerida.

The Endocerida and Tarphycerida were the major contributors to the cephalopod faunas of the Middle and Late Canadian (L Ord) and significantly to those of the Middle and Late Ordovician. Middle and Late Ordovian diversification adds the Actinocerida, the Oncocrida, and the Discosorida. The Oncocerida have their origin in the Basslerocertidae and therefor possibly the Tarphycerida, if not directly the Ellesmerocerida. The Discosorida likely have their origin back in the Plectronocerida, although no obvious link has been found. The origin of the Actinocerida has already been discussed.


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  • Chen and Teichert; Cambrian Cephalopods; Geology v.11, pp 647 -750, no.11 Nov 1983.
  • Flower, R.H.; The Nautiloid Order Ellesmeroceratida (Cephalopoda); Memoir 12, New Mexico Bureaur of Mines and Mineral Resources, Socorro NM, 1964
  • Furnish, W.M. and Glenister, Brain F; Nautiloidea-Ellesmerocerida, in the Treatise on Invertebrate Paleontology Vol K, Nautiloidea; GSA and University of Kansas Press, C.Teichert and R.C. Moore Eds, 1964.
  • Kroger, B. and Landing, E.; The earliest Ordovician cephalopods of Eastern Laurentia - Ellesmerocerids of the Tribes Hill Formation, Eastern New York. Journal of Paleontology v. 81; pp 841 - 857. Sept 2007
  • ________ Onset of the Ordovician cephalopod radiation – evidence from the Rochdale Formation (middle Early Ordovician, Stairsian) in eastern New York; Geological Magazine, Cambridge University Press, 2008
  • Teichert, C; Main Features of Cephalopod Evolution, in The Mollusca, v.12, Paleontology and neontology of Cephalopods, M.R. Clarke and E.R. Trueman, Eds, Academic Press, 1988
  • Wade, M; Nautiloids and their descendants: cephalopod classification in 1986; in Memoir 44, New Mexico Bureaur of Mines and Mineral Resources, Socorro NM, 1988.
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