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Jump to: ,
Eukarya Phylogeny



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Our original concept was to adopt a organization, similar to our approach to the Vertebrates. Unfortunately, the study of the basal Eukarya ("Protistology"), is at some distance yet from the kind of phylogenetic certainty that prevails among the vertebrates. Nevertheless, after 18 months of experimentation, we're now convinced that our current "best-guess cladogram" is fairly stable. So, with considerable trepidation, we're trotting out this tree for testing. In simplified, form, it proceeds as follows:

EUKARYA
   |--
   `--+--
      `--+--
         `--""
            |--+--
            |  |  |--
            |  |  `--
            |  `--
            |     |--
            |     `--
            `--+--
               `--
                  |--
                  `--

The derivation of this arrangement is discussed at Top Level Cladograms.

The uniting plants and animals doesn't seem to have a name, so we have given it one, "Metabiotiformes," for convenience. Until more of the are filled in, your best bet will probably be to consult the alphabetical index of taxa.

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The following represents an alternative phylogeny, based largely on Cavalier-Smith (2002). Though at first glance it may appear quite different from the previous tree, a closer look will show that it mostly differs only in the position of the root - between bikonts (eukaryotes with flagella in pairs) and unikonts (eukaryotes with singleton flagella), rather than within ( + ). Because there are rather a lot of them, I've listed taxa incertae sedis after the tree rather than within it.

Eukaryota (see below tree for synonyms)
   |--Unikonta
   |    |--
   |    `-- (includes  and )
   `--Bikonta (see below for synonymy)
        |  i. s.: Phagodinium [Phagodinida]
        |--
        `--+-- (includes , ,  and )
           `--

Bikonta [Anisomonadea, Anisomonadia, Biciliata, Cabozoa, Cyathobodonea, Eurymonada, Extrumonada, Histionida, Isomita, Jakobidea, Kinetomonada, Kinetomonadea, Phytomastigophorea, Proterozoa, Thecomonadia, Varicristia]

Eukaryota [Actinopodea, Alternatia, Archaeprotista, Archimycetes, Autotractea, Cnidosporidia, Cyclia, Eosarcodina, Fungilli, Gymnomycota, Gymnomyxa, Heliozoa, Heliozoia, Heterozooflagellata, Histionia, Hydraulea, Mastigophora, Metakaryota, Mycetozoia, Myxothallophyta, Neomonada, Neosarcodina, Neosporidia, Neozoa, Panstomata, Phycomycetes, Plasmodroma, Protista, Protoctista, Protomycota, Protoneozoa, Protophyta, Protoplasta, Protozoa, Rhizoflagellata, Rhizomastigida, Rhizopoda, Sarcodina, Sarcomastigophora, Sarcomastigota, Sarcomonadea, Zoomastigina, Zoomastigophorea]

Eukaryota incertae sedis:
 Melanocyrillium
 Trachyhystrichosphaera
 
 Haemogregarina aeglefini
 Haemohormidium terraenovae
 Mycelites ossifragus
 Pseudodevescovina uniflagellata
 Plagiopyxis
   |--P. callida
   `--P. declivis
 Dictyoporus
 Corollolithion kennedyi
 Helenea chiastia
 Lithraphidites acutus
 Rhagodiscus asper
 Marthasterites Deflandre 1959
   `--M. furcatus
 Eiffelithus
   |--E. eximius
   `--E. turriseiffelli
 Lithastrinus septenarius
 Eprolithus
   |--E. moratus
   `--E. octopetalus
 Micula decussata
 Quadrum gartneri
 Calculites anfractus
 Reinhardites antophorus
 Lucianorhabdus cayeuxii
 Lithocodium aggregatum
 Gonium
   |--G. pectorale
   `--G. sociale
 Cinetidomyxa chattoni
 Discobotellina biperforata
 Lagenidiopsis Golemansky 1974
   |--L. ancora Sudzuki 1979
   |--L. elegans (Gruber 1884)
   `--L. valkanovi Golemansky 1974
 Pseudolagenidiopsis Sudzuki 1979
   `--P. golemanskyi Sudzuki 1979
 Voluta Chardez 1972
   `--V. hemispiralis Chardez 1972
 Pseudovolutella Sudzuki 1979
   `--P. iriomotensis Sudzuki 1979
 Trichosporon cutaneum
 Spirophyton
 Bicorium Maslov 1956
   `--*B. kusbassense Maslov 1956
 Archaeolithothamnium
   |--A. irinae Maslov 1956
   `--A. saipanense [=Sporolithon saipanense]
 Wengania Zhang 1989
   |--*W. globosa Zhang 1989
   |--W. exquisita Zhang, Yin et al. 1998
   `--W. minuta Xiao 2004
 Paratetraphycus giganteus Zhang 1985
 Meghystricosphaeridium Zhang, Yin et al. 1998
   |--*M. chadianensis (Chen & Liu 1986) (see below for synonymy)
   |--M. densum (Kolosova 1991) Zhang, Yin et al. 1998
   |--M. gracilentum (Yin in Yin & Liu 1988) Zhang, Yin et al. 1998
   |--M. magnificum Zhang, Yin et al. 1998
   |--M. perfectum (Kolosova 1991) Zhang, Yin et al. 1998
   `--M. reticulatum
 Dicrospinasphaera Yuan & Hofmann 1998
   `--*D. zhangii Yuan & Hofmann 1998
 Apodostoides basileus Zhang et al. 1998
 Tianzhushania spinosa Yin & Li 1978
 Ericiasphaera
   |--E. magna (Zhang 1984)
   |--E. rigida
   `--E. sparsa
 Asterocapsoides sinensis
 Cerionopora ordinata [incl. Bacatisphaera baokangensis]
 Briareus borealis Knoll 1992
 Castaneasphaera speciosa
 Cymatiosphaeroides yinii
 Distosphaera speciosa Zhang et al. 1998
 Echinosphaeridium maximum
 Eotylotopalla
   |--E. dactylos
   `--E. delicata
 Filisphaeridium
 Goniosphaeridium
   |--G. acuminatum
   |--G. conoideum
   `--G. cratum
 Leiosphaeridia tenuissima
 Lophosphaeridium
 Mastosphaera changyangensis
 Micrhystridium
 Papillomembrana compta
 Pustulisphaera membranacea
 Sinosphaera rupina
 Vulcanisphaera phacelosa
 Gremiphyca corymbiata Zhang et al. 1998
 Paramecia incognata Zhang & Yuan 1992
 Thallophycoides phloeatus Zhang & Yuan 1992
 Monas
   |--M. minima
   |--M. socianilis
   `--M. stigmatica
 Iodamoeba butschlii
 Steatonyssus murinus
 Osagia monolamellosa
 Radiosus radialis
 Granomarginata
   |--G. dhalli
   |--G. primitiva
   `--G. simlaensis
 Griffonia simplicifolia
 Mizzia
   |--M. velebitana Schubert 1908
   `--M. yabei
 Pseudoepimastopora likana Kochansky-Devidé & Herak 1960
 Arabicodium [incl. Siamporidium Endo 1969]
 Halimeda soltanensis Poncet 1989
 Ortonella
   |--O. latifibrosa Endo in Endo & Kanuma 1954
   `--O. morikawai Endo 1958
 Nanopora
   |--‘Anthracoporella’ fragilissima Maslov 1939
   `--‘Anthracoporella’ kasachiensis Maslov 1939
 Roquesselsia Termier et al. 1977
   `--‘Anthracoporella’ insolita Ivanova 1988
 Oligoporella Pia 1912
 Clavaphysoporella Endo 1958
 Clavaporella [incl. Eoclypeina Poncet 1987 non Emberger in Vachard 1985]
   `--C. reinae Racz 1966 [incl. *Eoclypeina crassa Poncet 1987]
 Neoanchicodium Endo in Endo & Kanuma 1954
 Nostocites Maslov 1929
   `--*N. vesiculosa Maslov 1929
 
 Sphairionia Tien 1989 [incl. Pseudosphairionia Pronina 1996]
   |--*S. sikuoides Tien 1989 (see below for synonymy)
   `--S. gallinae Vachard, Hauser et al. 2001
 Tuberitinidae [Tuberitinaceae]
   |--Tuberitina collosa Reitlinger 1950
   |--Diplosphaerina Derville 1931
   |    `--D. inaequalis (Derville 1931)
   `--Eotuberitina Miklukho-Maclay 1958
 Marpolia spissa
 Chamaebotrys
   |--C. boergesenii
   `--C. lomentariae
 Coccobotrys
   |--*C. verrucariae
   `--C. mucosus
 Hyalostachybotrys
   |--H. bisbyi
   `--H. sacchari
 Streptobotrys
   |--S. arisaematis
   |--S. caulophyllii
   `--S. streptothrix
 Daffrania biloba
 Polysphaerinella bulla
 Luffisphaera [Holosea]
 Kamaena
   |--K. delicata
   `--K. tatarstanica
 Izhella nubiformis
 Shuguria flabelliformis
 Bosea Srivastava 1975
 Evittia Brito 1967
 Anthosphaeridium Cookson & Eisenack 1968
 Archaeodinium Shu 1979
 Artemisiocysta Benedek 1972
 Ascostomocystis Drugg & Loeblich 1967
 Beringiella Bujak 1984
 Caddasphaera Fenton et al. 1980
 Catillopsis Drugg 1970
 Ceratocystidiopsis Deflandre 1937
 Chonesphaera Klumpp 1953
 Cirrifera Cookson & Eisenack 1960
 Collumosphaera Jain & Dutta in Dutta & Jain 1980
 Conicoidium Jiabo 1978
 Cupritia Courtinat & Gaillard 1980
 Cyclodictyon Cookson & Eisenack 1958
 Cyclopsiella Drugg & Loeblich 1967
 Cystidiopsis Nagy 1965
 Deflandridium Nagy 1969
 Dictyosphaeridium Wetzel 1952
 Diplofusa Cookson & Eisenack 1960
 Wallodinium Loeblich & Loeblich 1968 [=Diplotesta Cookson & Eisenack 1960 (preoc.)]
 Domasiella Eisenack 1969
 Enigmadinium Wrenn & Hart 1988
 Epicephalopyxis Deflandre 1935
 Eyrea Cookson & Eisenack 1970
 Fromea Cookson & Eisenack 1958
 Halodinium Bujak 1984
 Halophoridia Cookson & Eisenack 1962
 Korojonia Cookson & Eisenack 1958
 Lophodictyotidium Pocock 1972
 Mathurosphaera Varma & Dangwal 1964
 Membranosphaera Samoilovich in Samoilovich & Mtchedlishvili 1961
 Mirificysta Yu 1982
 Nummus Morgan 1975
 Ollula Góczán 1962
 Palaeohystrichosphaeridium Eiserhardt 1986
 Palaeostomocystis Deflandre 1937
 Palhistiodinia Deflandre 1939
 Paralecaniella Cookson & Eisenack 1970
 Pronoctiluca Fabre-Domergue 1889 (see below for synonymy)
 Piperodinium Mudie in Scott et al. 1984
 Platycystidia Cookson & Eisenack 1960
 Polyasterias Meunier 1910
 Prismatocystis Habib 1969
 Pseudofromea Burger 1980
 Pterocystidiopsis Deflandre 1937
 Quadrina Bujak 1980
 Rhombodella Cookson & Eisenack 1962
 Schizocystia Cookson & Eisenack 1962
 Sicyoideum He 1980
 Spicadinium Batten & Lister 1988
 Spinuliferites Huang 1981
 Stylodiniopsis Eisenack 1954
 Tetragonidium Pascher 1914
 Tubulifera Schumacker-Lambry 1978
 Sinosabellidites Zheng 1980 [incl. Huainanella Wang 1982]
   `--*S. huainanensis Zheng 1980 [incl. Huainanella cylindrica Wang 1982]
 Vindhyania Mathur 1983
 Phimopollenites pannosus (Dettmann & Playford 1968) Dettmann 1974
 Sarjeantia Horowitz 1975
 Warburgella rugolosa
   |--W. r. rugolosa
   `--W. r. sinensis
 Prothyris lanceolata
 Ammoclathrinidae [Ammoconidae]
   |--Ammosolenia Haeckel 1889
   |    `--A. rhizammina Haeckel 1889
   |--Ammoclathrina Laubenfels 1948 [=Ammoconia Haeckel 1889 (preoc.)]
   |    |--*A. aulopegma (Haeckel 1889) [=Ammoconia aulopegma]
   |    `--A. sagenella (Haeckel 1889) [=Ammoconia sagenella]
   `--Ammolynthus Haeckel 1889
        |--*A. haliphysema Haeckel 1889
        `--A. prototypus Haeckel 1889
 Ophiocytium capitatum
   |--O. c. var. capitatum
   `--O. c. var. longispinum
 Cysticercus cellulosae
 Morania Walcott 1919
   |--*M. globosa Walcott 1919
   `--M. antiqua Fenton & Fenton 1939
 Longfengshania Du 1982
   `--*L. stipitata Du 1982
 Daltaenia Hofmann 1985
   `--*D. mackenziensis Hofmann 1985
 Globochaete Lombard 1945
 Ovummuridae
   |--Ovummurus Minoura & Chitoku 1979
   |    `--*O. duoportius Minoura & Chitoku 1979 [=O. duoportio]
   |--Minourella Munnecke, Servais & Vachard 2000
   |    `--*M. gotlandica Munnecke, Servais & Vachard 2000
   |--Arouxina Munnecke, Servais & Vachard 2000
   |    `--*A. pluricellata Munnecke, Servais & Vachard 2000
   `--Samtlebenella Munnecke, Servais & Vachard 2000
        `--*S. circumcamerata Munnecke, Servais & Vachard 2000
 Calcisphaeridae
   |--Calcisphaera Williamson 1880
   |--Polyderma Derville 1950
   |--Asterosphaerella Villain 1975
   `--Eovolvox Kazmierczak 1975
 Anisonema ovale
 Cyathomonas truncata
 Pleuromonas jaculans
 Thylacomonas compress
 Cyclopyxis arcelloides
 Nebula carinatalla
 Quadrulella
   |--Q. globulosa
   `--Q. symmetrica
 Subulamoeba saphirina
 Rosenia Waagen & Wentzel 1886
 Pseudodimorpha Mikrjukov 2000
   `--*P. marina (Fenchel et al. 1995) [=Tetradimorpha marina]
 Quadricilia rotunda (Skuja 1948)
 Moncharmontia apeninica
 Nummofallotia apula

*Meghystrichosphaeridium chadianensis (Chen & Liu 1986) [incl. M. acrotrichum Yin in Yin & Liu 1988, Tianzhushania tuberifera]

Pronoctiluca Fabre-Domergue 1889 [incl. Pelagorhynchus Pavillard 1917, Protodinifer Kofoid & Swezy 1921]

*Sphairionia sikuoides Tien 1989 [incl. Soroorina ghezelbashi Baghbani 1993 (n. n.), Sp. (*Pseudosphairionia) tienii Pronina 1996]

* Type species of generic name indicated

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