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Foraminifera are amoeboid protists (Kingdom Protista) that produce protective shells, also known as tests, which have openings or foramina for the extrusion of pseudopodia, from which the group derives its name. they are considered to comprise a taxonomic order, the Foraminiferida, within the rhyzopod subclass Granuloreticulosa.

Some classifiers have given them higher taxonomic rank, as high as phylum, which results in crowding of kingdom level taxa, and have substituted the Retaria for the Granuloreticulosa. Ranking foraminifera lower than order results in generic crowding and the need for extra subgenera and subspecies.

Foraminifera Biology

The living organism, loosely referred to as an animal, is unicellular or acellular, but not in the real sense of the word, primitive. Complex body functions performed by differentiated tissue and distinct organs in Metazoa are performed within the cell itself. As with Amoebida, the protoplasm in Foraminiferida is divided into an inner layer of darkish endoplasm surrounded by an outer layer of lighter ectoplasm. Pseudopoda, which are formed from ectoplasm, are anastomosing - branching and recombining. Endoplasm is confined to already constructed chambers, which are formed from ectoplasm and pseudopoda, and may be colored in shades of yellow, yellowish brown, greenish brown, salmon-rose, orange-red, or crimson.

Nuclei All forams have one or more nuclei, which are typically spherical. Nuclei of the more primitive, agglutinated or pseudochitinous genera are inflexible, enclosed in a thick membrane. Nuclei of more evolved forms, especially those with numerous narrow foramina, are more plastic, with thinner membranes.

Reproduction cycles Forams go through an alternation of generations in which an asexual stage with simple multiple fission (schizogeny) is followed by a sexual stage in which gametes are produced (gamogeny), so on and so forth. During the asexual phase the entire protoplasm is used , the parent being termed schizont or agamont, Resulting embryos are comparatively large and produce large proloculi.

Gametes During the sexual phase of reproduction gametes are produced which combine in pairs to form new individuals in which proloculi are small. Most forms studied so far are hologamic which produce unequally biflagellate gametes. In a few genera such as Allogromia and plastogamic forms such as Patelina and Rubratella, relatively large (40μ - 50μ in diameter) amoeboid gametes are produced. Other plastogamic genera such as Glabrotella have triflagellate gametes about 8μ in diameter. Biflagellate gametes typically vary in length from 2μ - 6μ and in width from 1.2μ - 3.5μ, Flagella range in length from 3μ for the smaller and 8μ for the larger to 5μ for the smaller and 20μ for the larger.

Pseudopodia Pseudopodia (informally pseudopods or false feet) in foraminifera are invariably granuloreticulose; composed of very elongate extensions of the protoplasm which readily bifurcate and anastomose. Individual pseudopods are only slightly thicker than the plasmatic granules streaming within them. Commonly they have a more firm axis surround by a more fluid layer. The relatively more solid axis and granular streaming are the most characteristic features.

The most important function of the pseudopods is in the capturing and digesting of prey, and in the expelling of debris. Other functions are in the construction of tests, forming of protective cysts, and making temporary or semipermanent attachment to substrate.

Foraminifera taxonomy

Recognized suborders from Loeblich & Tappan 1988, in more or less phylogenetic sequence:












The Carterinina, Robertinina, Lagenina, Globigerinina, and Spirillinina were removed form the Rotaliina in the Treatise on Invertebrate Paleontology Part C (Loeblich and Tappan 1964) where they were ranked as superfamilies; the Involutinina separated from the Cassidulinacea, the Silicoloculinina from the Litualacea (same).




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Cushman, Joseph A. (1950). Foraminifera, their classification and economic use, 4th Ed. Harvard University Press, Cambridge, Mass.

Loeblich, Alfred R Jr and Tappan H. Sarcodina, Chiefly "Thecamoebians and Foraminiferida. Treatise on Invertebrate Paleontology Part C, Protista 2. R.C. Moore (Ed). Geological Society of America and University of Kansas Press.



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BALB/c splenocytes were stained either with 5 M CFSE (Invitrogen) for 10 min at 37C and pulsed with the relevant AH-1 MHC class I peptide (corresponding to amino acids 423431 of gp70, SPSYVYHQF; CFSEhigh cells) or with 0.5 M CFSE and pulsed with an irrelevant peptide (HA) as a control (CFSElow cells). 107 splenocytes (per population) were transferred i.v. to each host. The draining inguinal lymph nodes and spleens were harvested 40 h later and analyzed by FACS. Percent lysis was calculated as 1 CFSEhigh/CFSElow normalized by the same ratio in naive BALB/c mice (%lysis = 100 x {1 [(CFSEhighexp/CFSElowexp)/(CFSEhighBALB/c/CFSElowBALB/c)]}). In previous research, the Wake Forest researchers engineered human bladders in the lab. In clinical trials, about 30 children and adults with congenital bladder abnormalities or bladder injuries who were treated with the engineered tissue showed normal or improved bladder function for nearly 10 years, Atala said. Other researchers have shown some success in clinical trials with transplanting windpipe tissue. Disclaimer: The authors of this article are responsible for its contents, including any clinical or treatment recommendations. No statement in this article should be construed as an official position of the Agency for Healthcare Research and Quality or the U.S. Department of Health and Human Services. If you experience ED and think that it may be a result of medication you are taking, do not stop taking the medication without first consulting your doctor. If the problem persists, your doctor may be able to prescribe a different medication. There are no formal tests to diagnose erectile dysfunction. Some blood tests are generally done to exclude underlying disease, such as diabetes, hypogonadism and prolactinoma. Apathy Drug and/or alcohol use Cialis works a bit faster (within about 15 minutes), and the effects last much longer -- up to 36 hours in some cases. Here Duplex ultrasound Dosing This is similar to magnetic resonance imaging. Magnetic resonance angiography uses magnetic fields and radio waves to provide detailed images of the blood vessels. Doctors may inject a "contrast agent" into the patient's bloodstream that causes vascular tissues to stand out against other tissues. The contrast agent provides for enhanced information regarding blood supply and vascular anomalies. Aside from the IV used to introduce the contrast material into the bloodstream, magnetic resonance angiography is noninvasive and painless. "The mechanisms underlying the association between RLS and erectile dysfunction could be caused by hypofunctioning of dopamine in the central nervous system, which is associated with both conditions," said Gao. Might surgery to treat impotence be necessary? Why or why not? Cialis May Help Ease Pulmonary Hypertension Treatment depends on the cause. Testosterone supplements may be used for cases due to hormonal deficiency. However, the cause is more usually lack of adequate penile blood supply as a result of damage to inner walls of blood vessels. This damage is more frequent in older men, and often associated with disease, in particular diabetes. The surgical procedure to implant the inflatable prosthesis is slightly more complicated than for a semi-rigid implant. Also, because there are more mechanical parts, there is a higher risk for mechanical failure requiring repair or adjustment. Thyroid function testsThyroid hormones regulate metabolism and the production of sex hormones; a deficiency may contribute to impotence. What is PMS? Unstable angina (chest pain due to coronary artery disease that occurs at rest or with minimal physical exertion)

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