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Gymnospermae ('naked seeds') include four classes, the , , and . They are that produce on an open (usually in a cone) rather than enclosed within a as in . This is a character for , and traditionally the gymnosperms have been thought to be with regard to the (flowering plants) with the as sister to angiosperms. However, recent molecular trees unexpectedly recover the gymnosperms as a group, as shown below. Cantino et al. (2007) suggested the use of the name "" for this clade, to distinguish it from the traditional usage of the name Gymnospermae. This has significant implications for our understanding of reproductive evolution in the seed plants. See Friedman and Floyd (2001) for a review of the alternative phylogenies. Notably, (previously believed a uniting character of Gnetales and angiosperms) may be more widespread among seed plants than previously believed, highlighting the potential dangers of extrapolating observations on a few exemplars to cover larger taxa.

A number of fossil taxa have been included in the seed plants in the past (see Doyle [1998] for a review), such as , , and . It is uncertain whether these fossil 'gymnosperms' would fall within a monophyletic clade of modern gymnosperms. The large gap in the fossil record between the earliest definite crown Acrogymnospermae (in the ) and the earliest angiosperms (in the early - Sun et al., 2002) suggests that at least some of the fossil 'gymnosperms' might be expected to lie on the angiosperm stem. In particular, and the have been nominated by different authors as angiosperm ancestors.

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Cantino, P. D., J. A. Doyle, S. W. Graham, W. S. Judd, R. G. Olmstead, D. E. Soltis, P. S. Soltis & M. J. Donoghue. 2007. Towards a phylogenetic nomenclature of Tracheophyta. Taxon 56 (3): E1-E44.

Doyle, J. A. 1998. Phylogeny of vascular plants. Annual Review of Ecology and Systematics 29: 567-599.

Friedman, W. E., & S. K. Floyd. 2001. Perspective: The origin of flowering plants and their reproductive biology – a tale of two phylogenies. Evolution 55 (2): 217-231.

Sun, G., Q. Ji, D. L. Dilcher, S. Zheng, K. C. Nixon & X. Wang. 2002. Archaefructaceae, a new basal angiosperm family. Science 296: 899-904.



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WebMD Feature Reviewed by Gary D. Vogin, MD About Erectile Dysfunction Click This test involves pumping fluid into the penis and measuring the pressure in the corpus cavernosum during an erection. This test will help establish how severe the venous leak is.Corpus Cavernosometry This is another test of venous leakage that involves infusing saline into the corpus cavernosum. The flow rate needed to maintain an erection indicates the degree of venous leakage. The leaking veins responsible may be visualised by infusing a mixture of saline and x ray contrast medium and performing a cavernosogram. SEXUAL HEALTH May cause shortening or numbness of the penis Erectile Dysfunction: Alternative Treatments Each supplement product differs a great deal in its exact formulation and the different ways in which it is produced. In general, male enhancement supplements increase erection effectiveness and staying power, improve sexual health, reduce premature ejaculation, increase orgasm power, boost libido, and restore youthful hormonal levels.The important point to remember is that not all products in the male enhancement industry are legitimate. There are lots of knock-off products. Only way to avoid such fake products is by educating yourself and through careful shopping. Prices of male enhancement products vary according to their manufactures. A reputable firm uses first-class ingredients and produce drugs in superior pharmaceutical-grade facilities, and thus their prices are very high compared to other brands. Build energy levels. In the same issue of the journal, Dr. Steven A. Grover and colleagues studied a group of 3,912 Canadian men, nearly half of whom reported having erectile dysfunction in the four weeks prior to visiting their family physicians. The men's cholesterol, glucose and blood pressure measurements were taken. What is premature ejaculation? Figure 1. Flow diagram of patient population in the study. This new study of Cialis, taken once daily, demonstrates that the drug substantially improves functional capacity, improves quality of life, and also results in less clinical worsening in patients with pulmonary arterial hypertension, Fonarow said. Erectile dysfunction. This inflatable penile prosthesis has 3 major components. The 2 cylinders are placed within the corpora cavernosa, a reservoir is placed beneath the rectus muscle, and the pump is placed in the scrotum. When the pump is squeezed, fluid from the reservoir is transferred into the 2 cylinders, producing a firm erection. Squeezing the top of the pump causes a reversal of flow of the fluid from the cylinders back into the reservoir. "If a man says 'no' to you with a lot of irritability, I still wouldn't take it personally or feel rejected, because it's very likely he's suffering some depression -- and again, it's not you," says Foley. Erectile dysfunction is treatable in all age groups. What Is the Penis? Sildenafil (Viagra), whose labeling was approved by the U.S. Food and Drug Administration (FDA) in March 1998, is the first oral medication to be marketed for the treatment of erectile dysfunction. Sildenafil inhibits the conversion of cGMP to guanosine monophosphate in the corpus cavernosum, thereby increasing the available concentration of cGMP. This reaction is largely catalyzed by the enzyme phosphodiesterase type 5. Sildenafil inhibits this enzyme, resulting in relaxation of the cavernosal smooth muscle.17 Sildenafil is absorbed rapidly, with peak plasma levels achieved within an hour. It undergoes hepatic metabolism and has a half-life of about four hours.

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