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Gymnospermae ('naked seeds') include four classes, the , , and . They are that produce on an open (usually in a cone) rather than enclosed within a as in . This is a character for , and traditionally the gymnosperms have been thought to be with regard to the (flowering plants) with the as sister to angiosperms. However, recent molecular trees unexpectedly recover the gymnosperms as a group, as shown below. Cantino et al. (2007) suggested the use of the name "" for this clade, to distinguish it from the traditional usage of the name Gymnospermae. This has significant implications for our understanding of reproductive evolution in the seed plants. See Friedman and Floyd (2001) for a review of the alternative phylogenies. Notably, (previously believed a uniting character of Gnetales and angiosperms) may be more widespread among seed plants than previously believed, highlighting the potential dangers of extrapolating observations on a few exemplars to cover larger taxa.

A number of fossil taxa have been included in the seed plants in the past (see Doyle [1998] for a review), such as , , and . It is uncertain whether these fossil 'gymnosperms' would fall within a monophyletic clade of modern gymnosperms. The large gap in the fossil record between the earliest definite crown Acrogymnospermae (in the ) and the earliest angiosperms (in the early - Sun et al., 2002) suggests that at least some of the fossil 'gymnosperms' might be expected to lie on the angiosperm stem. In particular, and the have been nominated by different authors as angiosperm ancestors.

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Cantino, P. D., J. A. Doyle, S. W. Graham, W. S. Judd, R. G. Olmstead, D. E. Soltis, P. S. Soltis & M. J. Donoghue. 2007. Towards a phylogenetic nomenclature of Tracheophyta. Taxon 56 (3): E1-E44.

Doyle, J. A. 1998. Phylogeny of vascular plants. Annual Review of Ecology and Systematics 29: 567-599.

Friedman, W. E., & S. K. Floyd. 2001. Perspective: The origin of flowering plants and their reproductive biology – a tale of two phylogenies. Evolution 55 (2): 217-231.

Sun, G., Q. Ji, D. L. Dilcher, S. Zheng, K. C. Nixon & X. Wang. 2002. Archaefructaceae, a new basal angiosperm family. Science 296: 899-904.

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