| Silurian period|
423 to 419 million years ago
Geography of the Ludlow
The geography of the Ludlovian world is best known for the on-going train wreck which began the assembly of Laurasia - or actually "Laurasia I" since there are arguably two continents by that name. By Ludlovian times, Baltica (including Scandinavia, European Russia and parts of Northern Europe) had collided with Laurentia (North America and Greenland). This began the north to south closure of the Iapetus Ocean which had previously separated the two. The western edge of Baltica was subducted under the Laurentian continent, with drastic effects in North America. These effects included the northern part of the Acadian Orogeny, a string of new volcanoes stretching from East Greenland (future home of Ichthyostega and other famous fossils) down the Laurentian coast towards New York and Pennsylvania.
In the meanwhile, the microcontinent of Avalonia, as well as bits and pieces of Southern Europe which had been flaking off the Gondwanan landmass, were running full tilt into the southern shores of Laurentia and Baltica. These fragments would add to what is now the lower eastern seaboard of the United States and Southern Europe, respectively. The exact timing of these events is not clear. Current thinking is that at least some of the European pieces arrived in Ludlovian times, but that Avalonia did not close the southern part of the Iapetus before the Pridoli, at the earliest.
Still further behind, Siberia was approaching the new Laurasian landmass from the northeast, followed by parts of what would eventually become Kazakhstan and Central Asia. The rest of the world looked much as it had since the beginning of the Cambrian, except that the entire Gondwanan landmass was continuing its slow slide across the South Pole which, in this era, passed through south central Africa. ATW
Ludlovian Climate and Environment
The Ludlow was a moderately warm era. The Ice Age which ended the Ordovician had long been over, although a small polar ice cap was probably still present in Africa. Oxygen had reached 50-100% of modern levels in the atmosphere, favoring the evolution of larger animals in suitable, shallow-water environments. There were a good many of these, since the Ludlow experienced the highest sea levels of the Paleozoic, and shallow seas covered many continental areas. Carbon dioxide levels were still quite high, perhaps 10 times today's concentrations. Indeed, carbon dioxide may actually have been increasing due to the extensive volcanic activity which accompanied the formation of Laurasia. The enormous Gondwanan peninsula formed by Australia and Antarctica prevented the formation of a continuous current around the South Pole, and the shores of the Gondwanan mainland were warmed by tropical waters flowing southeast through the Rheic Ocean. ATW
Dr David Gladwell of the University of Leicester describes a rare Silurian Lagerstätte:
- "An exceptionally preserved biota of Upper Silurian (Ludlow Series) age is found in Lower Leintwardine Formation Channel fill deposits around Leintwardine in the Central Welsh Borderland. There are six submarine channels in total, although only four outcrop and yield fossil faunas...The fauna is diverse, containing common representatives of Silurian biotas (such as brachiopods and trilobites), along with more unusual forms such as ophiuroid and asteroid seastars, eurypterids, xiphosurids and phyllocarids....(T)he echinoderms are mostly complete, whilst the majority of the arthropod material is made up of disarticulated components. Specimens are predominately preserved as hard-parts, although occasional soft-body preservation is encountered in the form of rare 'worm' fossils. In addition to the dominant invertebrate fauna, relatively rare disarticulated components of heterostracan fish are found; the sole taxon found, Archaegonaspis ludensis (Salter, 1859) is the earliest known British species..."
© David Gladwell, "An exceptionally preserved biota from Upper Silurian submarine channel deposits, Welsh Borderland" Annual Conference Abstracts - Palaeontological Association 46th Annual Meeting, Department of Earth Sciences, University of Cambridge, December 15-18, 2002
Even at this early date, the development of terrestrial ecosystems was well under way, as indicated by the following passage:
- "Sherwood-Pike and Gray (1985) describe probable ascomycete [this is unlikely! -- ATW] hyphae from Late Silurian (Ludlow) strata. They suggest that these were terrestrial Fungi at least contemporaneous with the earliest land plants. Some hyphae occurred inside small rounded pellets of apparent invertebrate fecal material; if this is correct, it suggests a fauna that included mycophagous (fungus-eating) microarthropods. Rolfe (1985b) accepts this suggestion and identifies millipedes from Late Silurian (Ludlow-Pridoli), now complemented by the discovery of predatory animals (Jeram et al. 1990). The importance these observations is that they confirm at least a minimal terrestrial food consisting of primary producers, decomposers, secondary consumers, and predators."
Anna K. Behrensmeyer et al, Terrestrial Ecosystems Through Time; © 1992 The University of Chicago
page by M. Alan Kazlev and ATW
page uploaded 10 June 2002
revised ATW030107 and MAK030427
(original uploaded on Kheper site 12 September 1998)
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