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Lycopodiopsida


Lycopsids, also known as lycophytes and lycopods, are a group of very ancient that are only distantly related to other land plants. They have a long history stretching back possibly to the late period. Their living representatives include club mosses and quillworts, but in past ages they dominate the landscape with huge forests.

Lycopods have roots, simple or branched stems and small, spirally arranged leaves (). The is either borne on a fertile leaf (a ) or is associated with one; it is thick-walled, and is either (producing only one kind of spore) or (producing two kinds of spore). The sperm cells are mobile and have two or many . are complex, with multicellular .

The earlier forms, such as from the late Silurian / early of , were small soft-bodied, low-growing plants which reached a typical height of 25 centimetres.

The group became especially successful and important during the late period, when the great formed huge swamp forests, with trees such as reaching over 40 metres in height.

The drying out of the climate during the latest Carboniferous was catastrophic for the lycopods, which require moist conditions to reproduce. All the giant "pole trees" like Lepidodendron died out, leaving only a number of small herbaceous types.

Today the lycopods are represented by five living genera with about 1100 species, all small creeping plants such as which reach only about 5 cm in height.

The following is a suggested classification

 Class Lycopsida ("club-mosses')
   Order  (ancestral types)
   Order 
   Order  (Club "mosses")
   Order  (Spike "mosses")
   Order  = Lepidocarpales (Scale trees) 
   Order 
   Order  (intermediate forms)
   Order  ("quillworts')

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Lycopodiopsida [Lepidophyta, Lycophyta, Lycophytae, Lycopsida, Microphyllophyta]
   |--Asteroxylon mackiei
   |--Baragwanathia
   `--Lycopodiophyta
        |-- [Lycopodiales]
        |    |--
        |    |--Huperzia
        |    |    |--H. lucidula
        |    |    `--H. selago (Linnaeus) Schrank & Mart. 1829
        |    |--Selaginellites Zeiller 1906
        |    |    |--*S. suissei
        |    |    `--S. hallei
        |    `--Phylloglossum drummondii
        `--

Lycopodiopsida incertae sedis:

 Lycopodites
 Lepidostrobus sternbergii [=Araucaria sternbergii, Araucarites sternbergii]
 Muscites
 Sigillaria Brongniart 1822 (see below for synonymy)
 Drepanophycus [Drepanophycaceae]
   `--D. spinaeformis
 Bacutriletes (van der Hammen) Potonié 1956
   |--*B. tylotus (Harris 1937) Potonié 1956
   |--B. guttula Archangelsky & Villar de Seoane 1991
   |--B. otwayensis Tosolini, McLoughlin & Drinnan 2002
   `--B. srivastavae Banerji, Jana & Maheshwari 1984
 Erlansonisporites Potonié 1956
   |--*E. erlansonii (Miner) Potonié 1956
   |--E. cerebrus Tosolini, McLoughlin & Drinnan 2002
   |--E. confertus Tosolini in McLoughlin et al. 2002
   |--E. decisum Tosolini, McLoughlin & Drinnan 2002
   `--E. indicus Banerji, Jana & Maheshwari 1984
 Hughesisporites Potonié 1956
   |--*H. galericulatus (Dijkstra) Potonié 1956
   |--H. australis Tosolini in McLoughlin et al. 2002
   |--H. coronatus Tosolini, McLoughlin & Drinnan 2002
   |--H. dettmanniae Tosolini, McLoughlin & Drinnan 2002
   |--H. rugulatus Archangelsky & Villar de Seoane 1991
   |--H. singhii Banerji, Jana & Maheshwari 1984
   `--H. variabilis Dettmann 1961
 Verrutriletes
   |--*V. compositipunctatus (Dijkstra) Potonié 1956
   |--V. depressus Tosolini, McLoughlin & Drinnan 2002
   |--V. dubius (Dijkstra) Potonié 1956
   `--V. tomtaylori Archangelsky & Villar de Seoane 1990
 *Ancorisporites binaensis Pant & Mishra 1986
 Banksisporites
   |--*B. pinguis (Harris) Dettmann 1961
   `--B. sinuosus Dettmann 1961
 Horstisporites Potonié 1956
   |--*H. reticuliferus (Dijkstra) Potonié 1956
   |--H. areolatus (Harris) Potonié 1956
   |--H. harrisii (Murray) Potonié 1956
   `--H. planatus (Marcinkiewicz) Marcinkiewicz 1971
 Favososporites Gamerro 1975
   |--*F. spinulosus Gamerro 1975
   `--F. brevis Tosolini in McLoughlin et al. 2002
 Henrisporites
   |--*H. affinis (Dijkstra) Potonié 1956
   |--H. musachioi Gamerro 1977
   |--H. selenacte (Douglas) Tosolini, McLoughlin & Drinnan 2002 (see below for synonymy)
   `--H. undulatus (Dijkstra) Potonié 1956
 Nathorstisporites hopliticus Jung 1958
 Maexisporites Potonié 1956
   |--*M. soldanellus (Dijkstra) Potonié 1956
   `--M. meditectatus (Reinhardt) Kannegieser & Kozur 1972
 Minerisporites
   |--*M. mirabilis (Miner) Potonié 1956
   |--M. laceratus Archangelsky & Villar de Seoane 1990
   |--M. marginatus
   `--M. patagonicus Archangelsky & Villar de Seoane 1989
 Paxillitriletes
   |--*P. reticulatus (Mädler) Hall & Nicholson 1973
   `--P. rintoulensis Tosolini in McLoughlin et al. 2002
 Ricinospora Bergad 1978
   `--*R. cryptoreticulata Bergad 1978
 Striatriletes van der Hammen ex Potonié 1956
   |--*S. sulcatus (Dijkstra) Potonié 1956
   `--S. imperfectus Tosolini in McLoughlin et al. 2002
 Tenellisporites Potonié 1956
   |--*T. tenellus (Dijkstra) Potonié 1956
   |--T. aachenensis (Dijkstra) Potonié 1956
   `--T. coronatus Baldoni & Batten 1991
 Trikonia Pant & Basu 1979
   |--*T. emarginata Pant & Basu 1979
   `--T. locmaniensis Tosolini in McLoughlin et al. 2002

Henrisporites selenacte (Douglas) Tosolini, McLoughlin & Drinnan 2002 [=Triletes selenacte Douglas 1973]

Sigillaria Brongniart 1822 (nom. cons.) non Rafinesque-Schmaltz 1819 [incl Rhytidolepis Sternb. 1821 (nom. rej.); Sigillariaceae]

* Type species of generic name indicated

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Alcock, F. J. 1938. Geology of Saint John region, New Brunswick. Geological Survey of Canada Memoir 216: 1-65.

Andersen, J. C. 1927. Popular names of New Zealand plants. Transactions and Proceedings of the New Zealand Institute 57: 905-977.

Barale, G., & M. Ouaja. 2002. La biodiversité végétale des gisements d’âge Jurassique supérieur-Crétacé inférieur de Merbah El Asfer (Sud-Tunisien). Cretaceous Research 23: 707-737.

Bateman, R. M., P. R. Crane, W. A. DiMichele, P. R. Kenrick, N. P. Rowe, T. Speck & W. E. Stein. 1998. Early evolution of land plants: phylogeny, physiology, and ecology of the primary terrestrial radiation. Annual Review of Ecology and Systematics 29: 263-292.

Cantino, P. D., J. A. Doyle, S. W. Graham, W. S. Judd, R. G. Olmstead, D. E. Soltis, P. S. Soltis & M. J. Donoghue. 2007. Towards a phylogenetic nomenclature of Tracheophyta. Taxon 56 (3): E1-E44.

Doyle, J. A. 1998. Phylogeny of vascular plants. Annual Review of Ecology and Systematics 29: 567-599.

Frey, W., M. Hofmann & H. H. Hilger. 2001. The gametophyte-sporophyte junction: Unequivocal hints for two evolutionary lines of archegoniate land plants. Flora 196: 431-445.

Gibson, N., & G. J. Keighery. 2000. Flora and vegetation of the Byenup-Muir reserve system, south-west Western Australia. CALMScience 3 (3): 323-402.

Iwatsuki, K. 1988. An enumeration of the pteridophytes of Nepal. In The Himalayan Plants vol. 1 (H. Ohba & S. B. Malla, eds.) The University Museum, University of Tokyo, Bulletin 31: 231-339.

Joseph, T. S., & C. B. Gena. 2002. A comparative study on the reproductive biology of three Indian species of Marsilea. Journal of the Bombay Natural History Society 99 (3): 583-586.

Kvaček, Z. 2002. Novelties on Doliostrobus (Doliostrobaceae), an extinct conifer genus of the European Palaeogene. Časopis Národního Muzea, Řada Přírodovědná 171 (1-4): 47-62.

Lötschert, W. 1954. Beitrag zur Pteridophyten-Flora von Mittel-Amerika. Senckenbergiana Biologica 35 (1-2): 109-119.

Ren, D., & J. Yin. 2003. New ‘osmylid-like’ fossil Neuroptera from the Middle Jurassic of Inner Mongolia, China. Journal of the New York Entomological Society 111 (1): 1-11.

Säilä, L. K. 2005. A new species of the sphenodontian reptile Clevosaurus from the Lower Jurassic of south Wales. Palaeontology 48 (4): 817-831.

Skog, J. E. 2003. Report of the Committee for Fossil Plants: 4. Taxon 52: 341.

Small, R. L., E. B. Lickey, J. Shaw & W. D. Hauk. 2005. Amplification of noncoding chloroplast DNA for phylogenetic studies in lycophytes and monilophytes with a comparative example of relative phylogenetic utility from Ophioglossaceae. Molecular Phylogenetics and Evolution 36: 509-522.

Tosolini, A.-M. P., S. McLoughlin & A. N. Drinnan. 2002. Early Cretaceous megaspore assemblages from southeastern Australia. Cretaceous Research 23: 807-844.

Yannitsaros, A. 1998. Additions to the flora of Kithira (Greece) I. Willdenowia 28: 77-94.

Zhang, R., & J. Pojeta, Jr. 1986. New bivalves from the Datang Stage, Lower Carboniferous, Guangdong Province, China. Journal of Paleontology 60 (3): 669-679.

Zherikhin, V. V. 2002. Ecological history of the terrestrial insects. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds.) pp. 331-388. Kluwer Academic Publishers: Dordrecht.

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