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The Nautiloidea are the basic which contain the ancestral orders as well as the modern . The Nautiloidea combines externally shelled cephalopods with rather simple septa that leave rather simple sutures but have siphuncles that are highly varied and often complex. This distinguishes them from the , the other ectocochliate cephalopods, in which the siphuncle is a simple featureless tube but the septa are complex, with complex and variable sutures.

Nautiloid shells vary from breviconic cyrtcones to orthconic longicones and from orthoconic brevicones to longiconic cyrtocones, that is are short, long, straight, curved, and may also be coiled in various, but ususally planospiral fashions.

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The in nautiloids, which is of prime importance in distinguishing orders and even families and genera, consists of segments that may be straight and tubular or expanded into the chambers in which the conncting rings may be thick and layered or thin and homogenous. Septal necks may be straight, outwardly curved, recumbent, or sometines inwardly sloped and may be short or long. Straight septal necks are orthochoanitic, outwardly curved ones are cyrtochoanitic. This contrasts with the prosiphonate siphuncles of the more advanced ammoinids wherein the necks point forward.

A unique feature found in many nautiloids are internal siphuncular deposits. These may be in the form of annular secretions, parietal linings, thick accumulations, or conical structures known as endocones. These features are found in the Endocerida, Actinocerida, Discosorida, Oncocerida, Orthocerida, and Pseudorithocerda and help distinguish them from each other and from other orders. Those lacking internal siphuncular deposits include the plectronocerids, ellesmerocerids, tarphycerids, many of the orthocerids and some of the oncocerids.

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The taxonomy presented here follows that of Mary Wade, 1988, in which related orders are combined in superorders. Note that the 'oidea' ending is used in the superordinal and not the subclassis sense. These are the

The Plectronoceratoidea includes the primitive ancestral and along with the and . The Nautilitoidea includes the , , and . The Orthoceratoidea includes the , , and .

The Endoceratoidea, Actinoceratoidea, and Discosoratoidea are each represented by a single order, respectively the Endocerida, Actinocerida, and Discosorida, and can be over looked as superfluous.

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Teichert, in the Treatise 1964, included the endocerids and actinocerids in separate subclasses, something Flower disagreed with, saying neither was any more distinct and diverse than any other order within the Nautiloidea. Similar treatment was not given the no less distinct and diverse Discosorida.

In Teichert's 1988 classification of the Cephalopoda the Nautiloidea as orginally and generally understood evaporates. Instead a number of more restricted subclasses appear. The Plectronocerida, Yanhecerida, Ellesmerocerida, and possible Protactinocerida, along with the Orthocerida and Ascocerida are combined in the Orthoceratoidea. Left out are the Pseudorthocerida. The Actinoceratoidea includes only the Actinocerida, as before, and the Endoceratoidea, the Endocerida and questionable Intejocerida. The Nautiloidea is reduced to include only the Tarphycerida, Discosorida, Oncocerida, and Nautilida (written in the publication as Nautiloidea) .

The Discosorida does not belong where placed, having a separate derivation and features not found in the other restricted nautiloid orders. Apart from the practical un-necessity of higher taxonomic terms, the Discosorida, Endocerida, and Actinocerida are all separately unique.

The Orthoceratoidea combines two distinct groups, the superorders Plectronoceratoidea and Orthoceratoidea of Wade, 1988; the latter being no less removed from the plectronoceroids than any other evolved group.

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  • Flower, R.H and Kummel, B; A Classification of the Nautiloidea; Journal of Paleontology, Vol 24, No.5, pp 604-616. Sept 1950
  • Flower, R.H.; The Nautiloid Order Ellesmeroceratida (Cephalopoda); Memoir 12, New Mexico Bureaur of Mines and Mineral Resources, Socorro NM, 1964
  • Teichert C, and Moore, R.C; Classification and Stratigraphic Distribution, in the Treatise on Invertebrate Paleontology Vol K, Nautiloidea; GSA and University of Kansas Press, C.Teichert and R.C. Moore Eds, 1964.
  • Teichert, C; Main Features of Cephalopod Evolution, in The Mollusca, v.12, Paleontology and neontology of Cephalopods, M.R. Clarke and E.R. Trueman, Eds, Academic Press, 1988
  • Wade, M; Nautiloids and their descendants: cephalopod classification in 1986; in Memoir 44, New Mexico Bureaur of Mines and Mineral Resources, Socorro NM, 1988.
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