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Panarthropods include true and their soft-bodied relatives, the and . These share a number of unique features (), such as " of a- which is molted regularly, appendages with chitinous claws, and mixocoel with metanephridia and ostiate heart (absent in tardigrades); several characters, such as the structure of the brain, indicate that arthropods and tardigrades are sister groups." Nielsen, "The Phylogenetic position of the Arthropoda"

During the period, panarthropods were much more diverse than today. There were several different lineages of and proto-arthropods, including large carnivorous swimming forms like Anomalocaris and Kerygmachela, as well as smaller crawling types.

Whilst true arthropods, tardigrades and onychophorans are highly differentiated in form today, such was not always the case. During the Early Cambrian, many transitional forms co-existed, and exciting new discoveries have come to light. Of particular interest is the afore-mentioned Kerygmachela, which provides a link between the lobopods, anomalocarids and arthropods. Another new anomalocarid-like taxon, Pambdelurion, indicates that the biramous arthropods actually arose from within the anomalocaridids. Budd, "Stem-group arthropods from the Lower Cambrian Sirius Passet fauna of North Greenland"

After dominating the oceans for many millions of years, the big protoarthropods died out with the end-Cambrian extinction, their place as super-predators being taken by during the .


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  • Claus Nielsen (1997). Animal Evolution: Interrelationships of the Living Phyla. Oxford University Press. .

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