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The Pennsylvanian Epoch of the Carboniferous Period
318 to 299 million years ago
The Pennsylvanian was the time of the great 'Coal Swamp Forests' which dominated the equatorial regions of the planet.

Lasting some 19 million or so years, the Late Carboniferous or Pennsylvanian was the high point of stem evolution, especially during the and . During this time the first and evolved and quickly diversified. By the end of the period these new forms, especially the synapsids, had supplanted the stem tetrapods as the dominant life form on land.

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During the late Carboniferous period and collide to form ; meanwhile comes up from the south. The resulting Appalachian, Ouachita, Marathon, Ural, Variscan, and Hercynian orogenies formed some of the largest mountains of all time. As a result of the collision of Gondwana and Laurasia the supercontinent of comes into being.

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On land, great forest swamps covered extensive equatorial areas. These forests consisted of diverse plants including tree ferns, which grew 15 meters in height, Calamites, a giant version of the modern "horsetail" plant, (e.g. , which attained a height of 30 metres), the extinct group of plants called "seed ferns", and primitive -like plants (Cordaites) that reached 40 meters in height

The extensive burial of biologically-produced carbon led to a buildup of surplus oxygen in the atmosphere; estimates place the peak oxygen content as high as 35%, compared to 21% today. This oxygen level resulted in insect and amphibian gigantism--creatures whose size is constrained by respiratory systems that are limited in their ability to diffuse oxygen. In this moist oxygen rich atmosphere flying insects were abundant, and some attained huge size, such as , with a wing span of 70 centimetres.

were abundant, especially the "labyrinthodonts," so called because of the complex (labyrinthine) pattern of folded in their teeth. They filled every available ecological niche, from fully aquatic eel-like forms, to large semiaquatic crocodile-like animals and small forms like modern day newts and salamanders, to terrestrial types similar to reptiles. Some types (the ) lost their legs altogether, superficially resembling snakes.

The earliest also evolved at this time, such as Hylonomus but remained relatively insignificant until the end of the period. Reptiles have a big advantage over stem tetrapods in that they do not have to return to water to breed; they can lay their eggs on dry land. So it is likely that with the appearance of reptiles the tetrapods were able to colonize the uplands for the first time, where they fed on an abundance of insects.

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The name "Carboniferous" derives from the fact that most of the important coal producing strata are of this age. However, it is specifically in the Late Carboniferous or Pennsylvanian sub-period that this is so. During this time most of the world's coal deposits were laid down, the coal being formed from compressed layers of rotting vegetation.

The large coal deposits of the Carboniferous primarily owe their existence to two factors. The first of these is the appearance of bark-bearing trees (and in particular the evolution of the bark fiber lignin). The second is the lower sea levels that occurred during the Carboniferous as compared to the Devonian period. This allowed for the development of extensive lowland swamps and forests in North America and Europe. Large quantities of wood were buried during this period because animals and decomposing bacteria had not yet evolved that could effectively digest the new lignin. Those early plants made extensive use of lignin. They had bark to wood ratios of 8 to 1, and even as high as 20 to 1. This compares to modern values less than 1 to 4. This bark, which must have been used as support as well as protection, probably had 38% to 58% lignin. Lignin is insoluble, too large to pass through cell walls, too heterogeneous for specific enzymes, and toxic, so that few organisms other than Basidiomycetes fungi can degrade it. It can not be oxidized in an atmosphere of less than 5% oxygen. It can linger in soil for thousands of years and inhibits decay of other substances. Probably the reason for its high percentages is protection from insect herbivory in a world containing very effective insect herbivores, but nothing remotely as effective as modern insectivores and probably many fewer poisons than currently. In any case coal measures could easily have made thick deposits on well drained soils as well as swamps.



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Optimal combination of drugs not known Related Health News The effectiveness of Enzyte is in dispute. Some medical professionals in fact advise against taking Enzyte, saying that it can lead to damage. The Center for Science in the Public Interest have urged the Federal Trade Commission to disallow further television advertising for Enzyte due to a lack of proper studies supporting claims. Ironically, while sex may be the last thing on your mind, it may help relieve PMS symptoms. Sexual activity stimulates the production of oxytocin and loads of feel-good endorphins that lower sensations to pain caused by premenstrual syndrome. Also, orgasms release the hormone dehydroepiandrosterone, or DHEA, which improves cognition and acts as an antidepressant. Erectile dysfunction can be defined as the inability to achieve or maintain a penile erection sufficient for satisfactory sexual performance.1 In the face of an aging population, decreasing social stigma associated with erectile dysfunction and an increasing availability of effective oral therapy for its treatment, the number of patients presenting with this complaint has increased dramatically. Current estimates describe 2–3 million Canadian men as having significant recurring erectile difficulties. Recent reports identifying a dramatic increase in rates of diabetes, increased longevity and higher quality-of-life expectations by "baby boomers" are all believed to be factors in a projected continued expansion of the patient population requesting medical help with sexual issues in the near future.2,3 Although historically erectile dysfunction was a problem identified and treated by urologists, today primary care physicians and other specialists write 80% of the prescriptions for sildenafil, the most popular drug used to treat the condition.4 Enzyte is said to contain: Tribulus terrestris; Yohimbe Extract; Niacin; Epimedium; Avena sativa; Zinc Oxide; Maca; Muira Pauma; Ginkgo biloba; L-Arginine; Saw Palmetto. Other ingredients: gelatin, rice bran, oat fiber, magnesium stearate, silicon dioxide. A Little Padding Helps, Too U.S. and Japanese researchers are investigating whether firing shock-waves towards the heart will help it sprout healthy new blood vessels. As many other medicament drug Levetra has side effects. The main Levetras’s side effects are following: Warning "There is an association between these two conditions, but we still don't know the exact mechanism," Gao said. His team said further research is warranted. Have you been able to achieve and maintain erections in the past? viagra for sale online An estimated 10 million to 20 million American men have some degree of erectile dysfunction, the incidence of which increases markedly with age. Between the ages of 40 and 70 years, the probability of complete erectile dysfunction triples from 5.1 percent to 15 percent.1 Previous SectionNext Section Have there been any recent changes in your life? DONALD W. SHENENBERGER, LT, MC, USNR, and Daily Once or twice a week A couple of times a month Viagra Helps COPD Patients Control Pulmonary Blood Pressure

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