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Jump to: ,
Perissodactyla


Burchell's Zebra, Equus quagga burchellii (, , )


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The Perissodactyla or odd-toed ungulates are and distinguished by having an odd number of toes on the (either 1, 3, or 5). Although early forms (Late through to early or middle ) were quite small, a number of lineages quickly evolved large to very large size by the middle Eocene. They were the primary mammalian herbivores from the Eocene to the , at which time they were supplanted by the . In contrast to the artiodactyls, perissodactyls are hindgut fermentors (that is, they have relatively simple stomachs supplemented by a large caecum to hold and ferment digested food) and a generally large middle toe.


(The following menu and phylogeny refers to subtopics of this page)


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  • :
  • Suborder:
`--viagra order no prescription
   |--
   |  `== ()
   |     `--+--
   |        `--
   `-- [Equoidea]
      |---
      |  `--
      |--
      `--+--
         `--


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:

  • Kingdom:
    • Phylum:
      • Class:
        • Order: Perissodactyla

Author: , 1848

Stratigraphic Range: Late to

Ecological role: and

The following sections are slightly modified from :

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The odd-toed arose in what is now in the late , less than 10 million years after the , in which the died out. By the start of the (55 million years ago) they had diversified and spread out to occupy several continents. The and both evolved in North America; the appear to have developed in from tapir-like animals and then reinvaded the Americas during the middle Eocene (about 45 million years ago). There were 12 families, of which only three survive. These families were very diverse in form and size; they included the enormous and the bizarre . The largest perissodactyl, an Asian rhinoceros called (also known as , and popularly known as ), reached 12 to 15 tons, more than twice the weight of an .

Perissodactyls were the dominant group of large terrestrial browsers right through the . However, the rise of grasses in the (about 20 million years ago) saw a major change: the with their more complex stomachs were better able to adapt to a coarse, low-nutrition diet, and soon rose to prominence. Nevertheless, many odd-toed species survived and prospered until the late (about 10,000 years ago) when they faced the pressure of human hunting and habitat change.

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The members of the order fall into two :

  • Hippomorpha are odd-toed ungulates that are fast runners with long legs. This suborder is represented by a few Eocene forms and the large and once diverse , which includes only one , Equus, comprising the horse, zebra, donkey, onager, and allied species. The Eocene family (members of which resemble more the Ceratomorpha) is traditionally included here as well, although its exact relationships are uncertain.
  • Ceratomorpha are odd-toed ungulates that have several functional toes and are heavier than and move slower than the Hippomorpha species. This suborder has two extant families: (tapirs) and (rhinoceroses)
the following section is from Palaeos com:

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Characters: elongate ; complete set of ; , often hypsodont & lophodont; horns, if present are unpaired on frontals or nasals & lack bone core; cervical vertebrae opisthocoelous; tall neural spines on ; absent; is long & narrow with small ; broad, tall with large gluteal (thigh) muscles; forelegs bear most weight, with hind legs providing propulsion; 3-4/3 (equids 1/1); "odd-toed" or mesaxonic, i.e. plane of symmetry passes through a large central digit; is primary weight-bearing ; , normally requiring constant foraging, but superior use (compared to ) of proteins and high-quality food materials.

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Perissodactyla
   |  i. s.: Helaletes
   |         Lophiodon
   |         Lophiaspis
   |         Symborodon
   |         Orotherium Marsh 1872
   |           |--O. cristonense Cope 1876
   |           |--O. laevii
   |           `--O. vintanum Marsh 1872
   |         Oligotomus
   |         Dolichorhinus
   |         Brontops
   |--Tapiromorpha
   |    |  i. s.: Heptodon tienshanensis
   |    |         Eomoropus
   |    |         Palaesyops
   |    |         Lambdotherium
   |    |--Orientolophus
   |    `--+--+--Cymbalophus Hooker 1984
   |       |  |    `--*C. cuniculus (Owen 1842) [=Hyracotherium cuniculus]
   |       |  `--Systemodon Cope 1881
   |       |       |--*S. tapirinus (Cope 1875) (see below for synonymy)
   |       |       `--S. etsagicus (Cope 1884) [=Hyracotherium angustidens etsagicum]
   |       `--+--Cardiolophus
   |          `--+--Homogalax protapirinus [=Systemodon protapirinus]
   |             `--+--Isectolophus
   |                `--+--
   |                   `--
   `--Hippomorpha [Equoidea]
        |  i. s.:  [Brontotherioidea]
        |           |--Metatelmatheriinae
        |           |    |--Qufutitan Wang & Wang 1997
        |           |    |    `--*Q. zhoui Wang & Wang 1997
        |           |    |--Metatelmatherium
        |           |    |    |--M. cristatum
        |           |    |    `--M. ultimum
        |           |    |--Desmatotitan
        |           |    |--Hyotitan
        |           |    |--Acrotitan
        |           |    `--Arctotitan
        |           |--Rhinotitan [Epimanteoceratinae]
        |           |    `--R. mongoliensis
        |           `--Embolotheriinae
        |--
        `--+--Hallensia mathewsi
           `--+--Hyracotherium Owen 1841
              |    |--*H. leporinum Owen 1841
              |    |--H. agile (Marsh 1873) [=Orohippus agilis]
              |    |--H. procyoninum (Cope 1872) [=Helotherium procyoninum]
              |    |--H. siderolithicum
              |    `--H. sylvaticum
              `--Palaeotheriidae
                   |  i. s.: Propachynolophus
                   |         Anchilophus
                   |         Plagiolophus
                   |--+--Palaeotherium
                   |  `--Propalaeotherium
                   `--Pachynolophus
                        |--P. lavocati
                        `--P. livinerensis

Systemodon tapirinus (Cope 1875) [=Hyracotherium tapirinum, Orohippus tapirinus; incl. H. cristatum Wortman 1896]

* Type species of generic name indicated

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Campbell, N. A. 1996. Biology (4th ed.) The Benjamin/Cummings Publishing Company, Inc.: Menlo Park (California).

Cope, E. D. 1876. Report upon the extinct Vertebrata obtained in New Mexico by parties of the expedition of 1874. Geographical Surveys West of the One Hundredth Meridian 4(2): i-iv, 1-370.

Froehlich, D. J. 2002. Quo vadis eohippus? The systematics and taxonomy of the early Eocene equids (Perissodactyla). Zoological Journal of the Linnean Society 134: 141-256.

Janis, C., & P. J. Jarman. 1984. The hoofed mammals. In All the World’s Animals: Hoofed Mammals (D. Macdonald, ed.) pp. 28-39. Torstar Books: New York.

Meng, J., Y. Hu & C. Li. 2003. The osteology of Rhombomylus (Mammalia, Glires): Implications for phylogeny and evolution of Glires. Bulletin of the American Museum of Natural History 275: 1-247.

Wang Y. & Wang J. 1997. A new brontothere from late Middle Eocene of Qufu, Shandong. Vertebrata PalAsiatica 35 (1): 68-77.

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Phylogeny: 01:15, 17 June 2008 (PDT) Characters and Links: ATW020617

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