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Jump to: ,
Parent taxa:
(check the following menu and phylogeny - the taxon in bold refers to the topic on this page)


Taxonomy Phylogeny

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         |-- [Harpoidea]
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Phacopida


from the of New York.


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This section from :

Phacopida is an of that lived from the to the . It is made up of a morphologically diverse group of related suborders.

The origin of the Phacopida order is uncertain. It comprises three suborders (, , and ) which share a distinctive (developmental stage lacking segments) type. The Cheirurina and Calymenina retain a plate (an ) but in virtually all Phacopina the free cheeks are yoked as a single piece. This sort of similarity in development suggests phylogenetic unity. The suborder Calymenina is the most primitive of the Phacopida order and shares some characteristics with the order , though it is not included in the subclass .

Phacopida had 8 to 19 thoracic segments and are distinguishable by the expanded , short or absent area, and (Phacopina) or holochroal (Cheirurina and Calymenina) eyes. Schizochroal eyes are compound eyes with up to around 700 separate lenses. Each lens has an individual which extended into a rather large .

The development of schizochroal eyes in phacopid trilobites is an example of post-displacement . The eyes of immature holochroal trilobites were basically miniature schizochroal eyes. In Phacopida, these were retained, via delayed growth of these immature structures (post-displacement), into the adult form.

and are two well-known represnetatives of this order. Other phacopids include , , and .


(The following menu and phylogeny refers to subtopics of this page)


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Taxonomy Phylogeny

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  • Phylum:
    • Class:
      • Order: Phacopida

Author: , 1864

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Phacopida
   |--Cheirurina
   |    |--Cheiruridae
   |    |    |  i. s.: Krattaspis Öpik 1937
   |    |    |--Eccoptochilinae
   |    |    |    |--Eccoptochile Hawle & Corda 1847
   |    |    |    |    |--*E. clavigera (Beyrich 1845)
   |    |    |    |    |--E. almadenensis Romano 1980
   |    |    |    |    |--E. impedita Hammann 1972
   |    |    |    |    |--E. mariana (Verneuil & Barrande 1855)
   |    |    |    |    |--E. perlata Hawle & Corda 1847
   |    |    |    |    |--E. quillieri (Tromelin in Guillier 1873)
   |    |    |    |    `--E. scrobiculata (Angelin 1854)
   |    |    |    |--Eccoptochiloides Prantl & Přibyl 1947
   |    |    |    |    |--E. scuticauda (Barrande 1846) [incl. *E. tumescens (Barrande 1852)]
   |    |    |    |    |--E. henryi Přibyl, Vaněk & Pek 1985
   |    |    |    |    `--E. octacantha (Angelin 1854)
   |    |    |    |--Foulonia Přibyl & Vaněk in Přibyl, Vaněk & Pek 1985
   |    |    |    |    `--*F. peregrina (Dean 1966) [=Ceraurinella peregrinus]
   |    |    |    `--Placoparina Whittard 1940
   |    |    |         `--*P. sedgwickii (McCoy 1849)
   |    |    |              |--P. s. sedgwickii
   |    |    |              `--P. s. shelvensis Hughes 1969
   |    |    |--
   |    |    `--+--
   |    |       `--
   |    `--Pliomeridae
   |         |  i. s.: Emsurella Rosova 1960
   |         |           `--E. humilla Rosova 1960
   |         `--Protopliomeropinae
   |              |--Protopliomerops Kobayashi 1934
   |              `--Rossaspis Harrington 1957
   `--Phacopina
        |--
        |--
        `--Greenops

* Type species of genus indicated

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Conway Morris, S. 1998. The Crucible of Creation. Oxford University Press: Oxford.

Crawford, R. L. 1992. Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History 8: 1-60.

Eckert, J. D., & C. E. Brett. 2001. Early Silurian (Llandovery) crinoids from the Lower Clinton Group, western New York State. Bulletins of American Paleontology 360: 1-88.

Fletcher, T. P. 2005. Holaspid variation in the solenopleurid trilobite Parasolenopleura gregaria (Billings, 1865) from the Cambrian of Newfoundland. Palaeontology 48 (5): 1075-1089.

Miller, S. A., & J. P. Harley. 1996. Zoology (3rd ed.) Wm. C. Brown Publishers: Dubuque (Iowa).

Owens, R. M. 2002. Cyclopygid trilobites from the Ordovician Builth-Llandrindod Inlier, central Wales. Palaeontology 45 (3): 469-485.

Přibyl, A., J. Vaněk & I. Pek. 1985. Phylogeny and taxonomy of family Cheiruridae (Trilobita). Acta Universitatis Palackianae Olomucensis Facultas Rerum Naturalium Geographica-Geologica XXIV 83: 107-193.

Prothero, D. R. 1998. Bringing Fossils to Life: An introduction to paleobiology. WCB McGraw-Hill: Boston.

Rong, J.-Y., R.-B. Zhan & J. Jin. 2004. The Late Odovician and Early Silurian pentameride brachiopod Holorhynchus Kiaer, 1902 from north China. Journal of Paleontology 78 (2): 287-299.

Sánchez, T. M., & N. E. Vaccari. 2003. Ucumariidae new family (Bivalvia, Anomalodesmata) and other bivalves from the Early Ordovician (Tremadocian) of northwestern Argentina. Ameghiniana 40 (3): 415-424.

Shu, D.-G., L. Chen, J. Han & X.-L. Zhang. 2001. An Early Cambrian tunicate from China. Nature 411: 472-473.

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