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Jump to: ,
Parent taxa:
(check the following menu and phylogeny - the taxon in bold refers to the topic on this page)


Taxonomy Phylogeny

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  • Superphylum:
  • Superphylum unspecified:
    • :
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    • :†
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Procoelomata



Among the many remarkable animals of the early Cambrian are those characterized by a protective covering of small hollow scales or sclerites. In most cases of course no trace of the animal is found, and the disarticulated sclerites are hard to associate with a particular organism. These tiny cap-shaped or scalelike skeletal elements make up the so called "" that are so predominant in assemblages of , and age (the so-called "").

Although the creatures that bore these scales remained a mystery for some time (and in many cases still do!), it is now known that many of these tiny shelly/skeletal elements belonged to small creeping slug-like animals. They formed a sort of interlocking "coat of mail", like medieval chain mail, that protected the animal against predators.

These animals cannot be easily classified in any modern phylum, but instead form a patchwork or of characteristics of several phyla, especially the , , and , regarding which they appear to be a sort of . would divide these creatures according to their modern relations (which may be a very difficult task, because often very little is known of them. In view of their close resemblance to each other however, another approach is to group them together in a single paraphyletic phylum, the "Procoelomata" of Bergström or the "Promollusca" of Kinman. Ironically, Bergström's term is probably too broad, since it would seem that these coat-of-mail animals are ancestral to the (the brachiopod-annelid-mollusc clade) only, rather than coelomate animals in general; whilst Kinman's term is certainly too narrow, since they were not solely mollusc cousins. However, I have decided to retain "Procoelomata" as this is the oldest term used to designate these creatures as a group.

Although most Procoelomata died out with the early Cambrian Botomian mass-extinction (one of the largest but least known extinction events in Earth's history), a few continued to thrive right up until the Devonian, and perhaps even the Carboniferous period

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Classification mostly according to Valid PMPD Orders.

It is not certain that all these groups are related; it has been suggested for example that the Machaeridia may be closer to the .


  • Class: Bengtson & Missarzhevsky, 1981
    • Order: Chancelloriida Walcott, 1920
    • Order: (incl. Halkieriida)
    • Order: Siphogonuchitida
  • Class: Cribricyathea
    • Order: Conoidocyathida
    • Order: Cribricyathida
    • Order: Vologdinophyllida
  • Class: Machaeridia
    • Order: Hercollepadida
    • Order: Turrilepadida (= Lepidocoleomorpha)
  • Class: Thylacocephala
    • Order: Concavicarida
    • Order: Conchyliocarida
  • Class: Tommotiida Missarzhevsky, 1970
    • Order: Mitrosagophora Bengtson, 1977
    • Order: Coleolida
    • Order: Hyolithelminthes
    • Order: Paiutiida
    • Order: Sabelliditida
    • Order: Volborthellida
  • Class and Order indet.
    • Family Anabaritidae Missarzhevsky, 1974

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  • Jan Bergström, 1989 "Metazoan evolution around the Precambrian-Cambrian transition", in The early evolution of Metazoa and the significance of problematic taxa, ed. by Alberto M. Simonetta and Simon Conway Morris, Cambridge University Press
  • Jan Bergström, 1989, The Origin of Animal Phyla and the New Phylum Procoelomata, Lethaia 22: 259-69.


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To bypass roadblocks along the brain-to-penis highway, drugs can be injected or inserted directly into the penis. While some men find this painful to think about, many others find that it can restore a satisfying sex life and is highly effective. Make an appointment to see a health professional within 1 to 2 weeks if an erection problem occurs more than 25% of the time and the problem: Questions to Ask Your Doctor about Impotence Inflatable implants consist of paired cylinders, which are surgically inserted inside the penis and can be expanded using pressurized fluid (see figure 3). Tubes connect the cylinders to a fluid reservoir and pump, which also are surgically implanted. The patient inflates the cylinders by pressing on the small pump, located under the skin in the scrotum. Inflatable implants can expand the length and width of the penis somewhat. They also leave the penis in a more natural state when not inflated. cialis without prescription Premature Ejaculation Surgery (radiation therapy, surgery of the colon, prostate, bladder, or rectum may damage the nerves and blood vessels involved in erection. Prostate and bladder cancer surgery often require removing tissue and nerves surrounding a tumor, which increases the risk for impotence.[8]) Unrealistic sexual expectations, which make sex a task rather than a pleasure Requires practice If unsuccessful, does not interfere with other treatments Sildenafil (Viagra), whose labeling was approved by the U.S. Food and Drug Administration (FDA) in March 1998, is the first oral medication to be marketed for the treatment of erectile dysfunction. Sildenafil inhibits the conversion of cGMP to guanosine monophosphate in the corpus cavernosum, thereby increasing the available concentration of cGMP. This reaction is largely catalyzed by the enzyme phosphodiesterase type 5. Sildenafil inhibits this enzyme, resulting in relaxation of the cavernosal smooth muscle.17 Sildenafil is absorbed rapidly, with peak plasma levels achieved within an hour. It undergoes hepatic metabolism and has a half-life of about four hours. A. MUSE (available since 1997) employs a small, specially-designed plastic plunger that is placed on the tip of the penis. Once the plunger is pressed, a rice-size pellet of medication (alprostadil) is pushed into the urethra. Moisture left by urine causes the pellet to dissolve, triggering an erection minutes later. Certainly less invasive than a hypodermic injection of medication, MUSE, an acronym for "medicated urethral system for erection," turns out to be a poor solution. Many men complain of a burning pain in the penis after the drug has been inserted, and there is a small risk of urethral injury. Since the efficacy of MUSE is extremely low - 10 percent or less, according to on 1998 study and the side effects are worse than injection therapy, I don't recommend it as a first-line treatment after a radical prostatectomy. Duplex ultrasound is used to evaluate blood flow, venous leak, signs of atherosclerosis, and scarring or calcification of erectile tissue. Injecting prostaglandin, a hormone-like stimulator produced in the body, induces erection. Ultrasound is then used to see vascular dilation and measure penile blood pressure. Measurements are compared to those taken when the penis is flaccid. Taking Viagra Won't Spur Risky Sexual Behavior But that may now all change with the concept of an alprostadil cream named Befar, (it may also be released later in some countries under the trade names of Alprox and Topiglan). This is the first topical cream in the world for the treatment of erectile dysfunction. Serfas, a manufacturer of biking accessories, set out to design a seat that would shift the rider's weight off the perineum. The result, called "the Eliminator," has a long groove down its middle and is hollowed out in front. In April 1999, researchers tested the newly designed seat on 15 regular cyclists, most of whom pedaled between 150 and 300 miles weekly. Decreased free T4 Hyperthyroidism Heat intolerance; weight loss; diaphoresis; palpitations Lid lag; exophthalmos; hyperreflexia; tremor; tachycardia Decreased TSH The workers studied did not have to spend years in the factory to develop problems -- sexual dysfunction began in new workers after just months on the job, Li said. Some insurance plans cover sex therapy. Once you select a therapist, call her/him to ask about whether or not his or her services are covered by your insurance provider. If payment cannot be worked out through insurance, many therapists can adjust their fees. Sex therapy clinics tend to be less expensive than private therapists. The cost of sex therapy programs varies, but generally range between $600 and $2,300 dollars, depending on the nature of the problem and the response to treatment.

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