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Systematics

  • Class
  • Subclass
  • Order Prolecanitida

The Prolecanitida are extinct cephalopods with discoidal to thinly lenticular shells and goniatitic or ceratitic sutures that retained the primitive retrochoanitic siphuncle. As typical for ammonoids the siphuncle is along the ventral margin. Prolecanitids form a relatively small and stable order within the Ammonidea with 43 named genera and about 1250 species. Prolecanitids have a stratigraphic range of about 108 m.y, stretching from the Late to the , and although not as numerous and diverse as their goniatitid contemporaries, they provided the stock from which all later Mesozoic ammonoids are derived.

The Origin of the Prolecanitida may be found in the , originally included in the but recently removed to the . Following their inception late in the Devonian, the Prolecanitida split into two lineages, idenfied by superfamilies; the earlier U Dev -M Perm in which the shells are fairly smooth and characteristicly have a large umbilicus and the later U Miss-Trias in which the umbilicus is small and there is moderate sculpture along the flanks. The Prolecanitaceae, through the derived family is the source for the Ceratitida, beginning in the late Permian with the ceratitid family .

Prolecanitids show long-term, gradual changes in shell geometry and the most limited use of available forms (euphamistcaly morphospace) as compared to the dominent goniatitds. Prolecanitid genera lasted an average of 14.7 m.y compared to 5.7 m.y. for Pennsylvanian (Upper Carboniferous) goniatitids.

While shell morphology was rather stable, with a limited number or morphologic options utilized, suture morphology in the Prolecanitida changed dramatically over time, from very simple sutures in the earliest (Devonian) genera to extremely complex-sutured genera in the late Paleozoic. The increase in suture complexity over their 108 m.y. duration resulted from the iterative of addition of as many as 12–15 replicate, U-shaped umbilical lobes, originating at the umbilicus and migrating across the flanks during both ontogeny and phylogeny, serration of lobes, and the subdivision of lateral and ventral lobes

Not only did the Prolecanitida evolve their sutures differently than in the Goniatitda, by increasing the number of umbilical lobes rather than by subdivision of the lateral saddle, their body chambers were short by comparison. This brings up the question of, just how closely related were the Prolecanitida and their Mesozoic descendants to the goniatitids in the first place.

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  • Prolecanitina in Miller, Furnish, and Schindewolf 1957. Paleozoic Ammonoidea; Treatise on Invertebrate Paleontology, Part L, pp L69 -L75
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