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These are the scorpions, carnivorous arachnids that have their pedipalps modified into powerful, claw-like pincers. All scorpions have the last six segments of their bodies, the metasoma, modified into a tail that bears the telson (the segment placed on the sixth segment), which has been modified into a venom-injecting organ used for subduing prey and potential predators.

Scorpions have an extensive, albeit notoriously patchy fossil record extending all the way back to the , where they began as a group of small, eurypterid-like creatures. They made the transition from marine to terrestrial environments some time between the late Silurian and early Devonian. All of the marine scorpions would eventually die out during the Devonian due to either climatic changes, or competition and or predation from various predatory and the larger, fiercer .

Scorpiones [Scorpiniata]
   |  i. s.: Belisarius xambeui Simon 1879 GE02
   |         Euscorpius [Euscorpiidae] L03
   |           |--E. flavicaudis L03
   |           `--E. italicus SF01
   |         Anuroctonus GE02
   |         Uroctonus GE02
   |         Vaejovis [Vaejovidae] AS-N08
   |           |--V. bilineatus Pocock 1898 AS-N08
   |           |--V. eusthenura SF01
   |           `--V. spinigerus (Wood 1863) AS-N08
   |         Mesophonus W70
   |          OA02
   |         Joctonus manicatus Thorell 1876 KK84
   |         Dacurus galbineus (Koch 1838) [=Centrurus galbineus, C. galbinus (l. c.), Caucon galbineus] K92
   |         Smeringurus aridus SF01
   |         Megacormus gertschi SF01
   |         Brotheas SF01
   |           |--B. amazonicus Lourenço 1988 L-M-HG08
   |           `--B. granulatus SF01
   |         Hemilychas alexandrinus (Hirst 1911) V05
   |         Broteochactas V-MDR07
   |           |--B. mapuera Lourenço 1988 L-M-HG08
   |           `--B. trezzi V-MDR07
   |         Stoermeroscorpio S07
   |         Prearcturus S07
   |         Pseudouroctonus apacheanus (Gertsch & Soleglad 1972) AS-N08
   |--Palaeoscorpius GE02
   `--+--+-- GE02
      |  `--Proscorpius [Palaeoscorpiones] GE02
      |       `--P. osborni GE05
      `--Protoscorpiones GE02
           |--Dolichophonus GE02
           `--Palaeophonidae GE02
                |--Palaeophonus GE02
                `--Allopalaeophonus GE02

* Type species of generic name indicated

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[AS-N08] Aguiar, A. P. N., P. L. Santana-Neto, J. R. B. Souza & C. M. R. de Albuquerque. 2008. Relationship between litter characteristics and female size in Tityus stigmurus (Scorpiones, Buthidae). Journal of Arachnology 36 (2): 464-467.

Giribet, G., G. D. Edgecombe, W. C. Wheeler & C. Babbitt. 2002. Phylogeny and systematic position of Opiliones: A combined analysis of chelicerate relationships using morphological and molecular data. Cladistics 18: 5-70.

[GE02] Giribet, G., G. D. Edgecombe, W. C. Wheeler & C. Babbitt. 2002. Phylogeny and systematic position of Opiliones: A combined analysis of chelicerate relationships using morphological and molecular data. Cladistics 18: 5-70.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[K92] Karsch, F. 1892. Arachniden von Ceylon und von Minikoy gesammelt von den Herren Doctoren P. und F. Sarasin. Berliner Entomologische Zeitschrift 36 (2): 267-310.

[KK84] Koch, L., & E. Keyserling. 1884-1890. Die Arachniden Australiens nach der Natur beschrieben und abgebildet vol. 2. Bauer & Raspe: Nürnberg.

[L-M-HG08] Lo-Man-Hung, N. F., T. A. Gardner, M. A. Ribeiro-Júnior, J. Barlow & A. B. Bonaldo. 2008. The value of primary, secondary, and plantation forests for Neotropical epigeic arachnids. Journal of Arachnology 36 (2): 394-401.

[L03] Lourenço, W. R. 2003. The first scorpion fossil from the Cretaceous amber of France. New implications for the phylogeny of Chactoidea. Comptes Rendus Palevol 2: 213-219.

[OA02] Ochoa, J. A., & L. E. Acosta. 2002. Orobothriurus atiquipa, a new bothriurid species (Scorpiones) from Lomas in southern Peru. Journal of Arachnology 30: 98-103.

[S07] Shultz, J. W. 2007. A phylogenetic analysis of the arachnid orders based on morphological characters. Zoological Journal of the Linnean Society 150 (2): 221-265.

[SF01] Soleglad, M. E., & V. Fet. 2001. Evolution of scorpion orthobothriotaxy: A cladistic approach. Euscorpius 1: 1-38.

[V-MDR07] Villarreal-Manzanilla, O., C. DoNascimiento & C. J. Rodríguez. 2007. Two new species of the enigmatic genus Stenophareus (Opiliones: Laniatores: Stygnidae) from the Venezuelan Guiana Shield. Zootaxa 1471: 43-51.

[V05] Volschenk, E. S. 2005. A new technique for examining surface morphosculpture of scorpions. Journal of Arachnology 33: 820-825.

[W70] Wills, L. J. 1970. The Triassic succession in the central Midlands in its regional setting. Quarterly Journal of the Geological Society of London 126: 225-283.


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