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Mostly with thick rigid walls containing and ; often forming . In contrast, one subclade, the , has lost the cell wall, and is mostly intracellular parasites or symbionts (Mycoplasma).

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The clade that I am here referring to as 'Clostridea' is probably more often seen under the name 'Firmicutes', and appears as such in the latest edition of Bergey's Manual of Systematic Bacteriology, probably the reference most often regarded as the one-stop shop for bacterial classification. However, the class 'Firmicutes', when first coined in Bergey's Manual, was originally intended to include all bacteria, including , and its restriction to one subset of that group strikes me as unfortunate. In contrast, the name 'Clostridea' that was used by Cavalier-Smith (1998) to refer to this taxon has not appeared in any other sense, and has the advantage of clearly referring to , one of the main genera within this clade. Admittedly, Cavalier-Smith excluded from Clostridea (Cavalier-Smith has no objections to the use of paraphyletic taxa in his classifications), but as the Mollicutes are probably embedded deep within this clade, and represent a relatively small part of it, I see no problem with expanding the name to cover them.

Perhaps more than any other group of bacteria, the taxonomy of Clostridea is currently unstable (and that's saying something!) In the past, members of Clostridea have generally been placed in a few large genera (such as and ), based on morphological/physiological grounds, that now appear to be highly polyphyletic. While new genera have been proposed to subdivide the old for a number of cases, researchers outside the bacterial taxonomy 'clique' have been slow to take them up. Many Clostridea are of significant ecomonomic importance, for instance medically (Clostridium tetani, Staphylococcus aureus, Streptococcus pneumoniae), in food production (Lactobacillus delbrueckii ssp. bulgaricus) or in biocontrol (Bacillus thuringiensis), and a significant amount of literature is already about under the old names for these taxa. Clostridea is therefore perhaps a perfect case study for the conflict that often exists between the creation of a rational, phylogeny-based classification and the researchers who have to actually try to use the resulting classification.

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Clostridea [Firmicutes, Teichobacteria]
   |  i. s.: Thermoacetogenium MoKi03
   |         Acetoanaerobium GH01a
   |         Oscillospira GH01a
   |         Syntrophococcus GH01a
   |--Thermoanaerobacteriaceae [Thermoanaerobacteriales] LK01
   |    |--Thermoanaerobacterium saccharolyticum GH01a, GH01b
   |    |--Ammonifex degensii GH01a, MiKo03
   |    |--Sporotomaculum GH01a
   |    |--Thermoanaerobium Zeikus, Hegge & Anderson 1979 GH01a, ZHA79
   |    |    |--*T. brockii Zeikus, Hegge & Anderson 1979 ZHA79
   |    |    `--T. aotearoense T01
   |    |--Thermoanaerobacter GH01a
   |    |    |--T. acetoethylicus GH01b
   |    |    |--T. brockii GH01b
   |    |    |--T. ethanolicus IT03
   |    |    `--T. tengcongensis CF04
   |    `--Coprothermobacter GH01a
   |         |--C. platensis GH01c
   |         `--C. proteolyticus GH01c
   `--+--Moorella thermacetica LK01, PK03 [=Clostridium thermaceticum PK03]
      `--+-- LK01
         |-- LK01
         `--Clostridiaceae [Clostridiales] LK01
              |-- GH01a
              |--Acetivibrio GH01a
              |--Acidaminobacter hydrogenoformans GH01a, PH03
              |--Anaerobacter GH01a
              |--Caloramator fervidus GH01a, PK03
              |--Natronincola GH01a
              |--Oxobacter pfennigii GH01a, PK03
              |--Sporobacter GH01a
              `--Thermobrachium GH01a

* Type species of generic name indicated

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Cavalier-Smith, T. 1998. A revised six-kingdom system of life. Biological Reviews 73: 203-266.

[CF04] Creevey, C. J., D. A. Fitzpatrick, G. K. Philip, R. J. Kinsella, M. J. O’Connell, M. M. Pentony, S. A. Travers, M. Wilkinson & J. O. McInerney. 2004. Does a tree-like phylogeny only exist at the tips in the prokaryotes? Proceedings of the Royal Society of London B 271: 2551-2558.

[GH01a] Garrity, G. M., & J. G. Holt. 2001a. The road map to the Manual. In Bergey’s Manual of Systematic Bacteriology, 2nd ed., vol. 1. The Archaea and the Deeply Branching and Phototrophic Bacteria (Boone, D. R., R. W. Castenholz & G. M. Garrity, eds) pp. 119-166. Springer.

[GH01b] Garrity, G. M., & J. G. Holt. 2001b. Phylum BIII. Thermodesulfobacteria phy. nov. In Bergey’s Manual of Systematic Bacteriology, 2nd ed., vol. 1. The Archaea and the Deeply Branching and Phototrophic Bacteria (Boone, D. R., R. W. Castenholz & G. M. Garrity, eds) pp. 389-393. Springer.

[GH01c] Garrity, G. M., & J. G. Holt. 2001c. Phylum BIX. Deferribacteres phy. nov. In Bergey’s Manual of Systematic Bacteriology, 2nd ed., vol. 1. The Archaea and the Deeply Branching and Phototrophic Bacteria (Boone, D. R., R. W. Castenholz & G. M. Garrity, eds) pp. 465-471. Springer.

[LK01] Ludwig, W., & H.-P. Klenk. 2001. Overview: A phylogenetic backbone and taxonomic framework for procaryotic systematics. In Bergey’s Manual of Systematic Bacteriology, 2nd ed., vol. 1. The Archaea and the Deeply Branching and Phototrophic Bacteria (Boone, D. R., R. W. Castenholz & G. M. Garrity, eds) pp. 49-65. Springer.

[MiKo03] Miroshnichenko, M. L., N. A. Kostrikina, N. A. Chernyh, N. V. Pimenov, T. P. Tourova, A. N. Antipov, S. Spring, E. Stackebrandt & E. A. Bonch-Osmolovskaya. 2003. Caldithrix abyssi gen. nov., sp. nov., a nitrate-reducing, thermophilic, anaerobic bacterium isolated from a Mid-Atlantic Ridge hydrothermal vent, represents a novel bacterial lineage. International Journal of Systematic and Evolutionary Microbiology 53: 323-329.

[MoKi03] Mori, K., H. Kim, T. Kakegawa & S. Hanada. 2003. A novel lineage of sulfate-reducing microorganisms: Thermodesulfobiaceae fam. nov., Thermodesulfobium narugense, gen. nov., sp. nov., a new thermophilic isolate from a hot spring. Extremophiles 7: 283-290.

[PK03] Parshina, S. N., R. Kleerebezem, J. L. Sanz, G. Lettinga, A. N. Nozhevnikova, N. A. Kostrikina, A. M. Lysenko & A. J. M. Stams. 2003. Soehningia saccharolytica gen. nov., sp. nov. and Clostridium amygdalinum sp. nov., two novel anaerobic, benzaldehyde-converting bacteria. International Journal of Systematic and Evolutionary Microbiology 53: 1791-1799.

[PH03] Pikuta, E. V., R. B. Hoover, A. K. Bej, D. Marsic, E. N. Detkova, W. B. Whitman & P. Krader. 2003. Tindallia californiensis sp. nov., a new anaerobic haloalkaliphilic, spore-forming acetogen isolated from Mono Lake in California. Extremophiles 7: 327-334.

[T01] Trüper, H. G. 2001. Etymology in nomenclature of procaryotes. In Bergey’s Manual of Systematic Bacteriology, 2nd ed., vol. 1. The Archaea and the Deeply Branching and Phototrophic Bacteria (Boone, D. R., R. W. Castenholz & G. M. Garrity, eds) pp. 89-99. Springer.

[ZHA79] Zeikus, J. G., P. W. Hegge & M. A. Anderson. 1979. Thermoanaerobium brockii gen. nov. and sp. nov., a new chemoorganotrophic, caldoactive, anaerobic bacterium. Archives of Microbiology 122: 41-48.


Characteristics and nomenclature CKT080123; phylogeny 09:31, 16 October 2010 (UTC)

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