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The Three Domains

The three-domain system or Three Domains is a biological classification introduced by in 1990 that emphasizes his separation of into two groups, originally called Eubacteria and Archaebacteria. Woese argued based on differences in genes that these two groups and the each arose separately from an ancestral progenote with poorly developed genetic machinery. To reflect these primary lines of descent, he treated each as a domain, divided into several different . The groups were also renamed the , , and , further emphasizing the separate identity of the two prokaryote groups.

A phylogenetic tree based on data, showing the separation of bacteria, archaea, and eukaryotes.

Although the three- system was quickly adopted by most molecular systematists, biologists like criticized him for over-emphasizing the uniqueness of the archaebacteria and ignoring strong genetic similarities between the groups. Subsequent studies have confirmed that the archaea are unusual in the composition of their cell membrane and structure of their . Other significant differences include archaeal systems for and which bear distinct similarity to those found in eukaryotes. For instance, archaeal consists of up to 14 subunits, whereas most bacterial RNA polymerases have only 4 subunits. Analysis of these subunits suggests that they are more closely related to those found in eukaryotes. Also, the archaea produce a number of DNA-binding proteins with similarity to eukaryotic histones.

Nevertheless, a minority viewpoint suggests retaining the older (Prokaryota and Eukaryota) and using the word bacterium in its earlier meaning of prokaryote.

Which system is preferable depends partly on the relationships of the organisms in question. Although the is discredited, molecular trees tend to group living things into the three domains, with the and the eubacteria forming a separate branch. However, it has been suggested this is an artifact of and that the root may instead belong among the eubacteria, in which case many eubacterial lines diverged before the archaebacteria did.

In 2006, the discovery of unique properties of the started a discussion about classification of (certain) viruses as fourth domain of life.


Credits: This article is licensed under the . It uses material from the ; copied to Palaeos org MAK090629

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Women and ED: Striking a Balance GORDON F. GREEN, CDR, MC, USNR Birth Control Patch Side Effects Vardenafil should not be used with alpha-blockers, medicines used to treat high blood pressure and benign prostate hypertrophy (BPH), because the combination of vardenafil and an alpha-blocker may lower the blood pressure greatly and lead to dizziness and fainting. Examples of alpha-blockers include tamsulosin (Flomax), terazosin (Hytrin), doxazosin (Cardura), alfuzosin (Uroxatrol), and prazosin (Minipress). Psychogenic impotence is relatively uncommon. It is characterized objectively by the presence of good nocturnal and morning erections and negative findings from all other tests. During the interview, a history of highly variable erections that can be totally absent one day but virtually normal the next suggests a psychogenic cause. Virtually 100% of men with severe depression have erectile dysfunction (ED). Sildenafil (Viagra) works well for psychogenic ED; other treatment modalities are also effective because the tissues, nerves, hormone levels, and vasculature are normal. The authors usually recommend a full psychological evaluation in these patients so that the underlying etiology can be identified and treated appropriately rather than just treating the symptom of ED. Therefore, the authors defer treatment of the patient's ED until he has begun psychological testing and therapy for the underlying problem. How Effective Is Vascular Reconstructive Surgery? Working with doctors to select medications that do not impair erectile function, Men taking erythromycin or ketoconazole should not take more than 5 mg of vardenafil in a 24-hour period. Potential Conflicts of Interest: Grants received: V. Snow (Bristol-Myers Squibb, Centers for Disease Control and Prevention, Novo Nordisk, Merck Vaccines, Boehringer Ingelheim, Wyeth, Sanofi Pasteur), P. Shekelle (Pfizer). Royalties: P. Shekelle (Up-to-Date). Any financial and nonfinancial conflict of interest of the group members were declared, discussed, and resolved. Avoid the use of illegal drugs. "A lot of men don't have physicians," Thompson explained. "They may not know what their blood pressure is or their lipid profiles, or they may be smokers, and they may never have been counseled to stop smoking or to reduce their weight. Twenty-four percent of those in the control group had erectile dysfunction. If so, it could offer a lifeline to patients who are too ill to undergo heart bypass surgery. Laboratory Tests These drugs should not be used with certain other medications, including nitroglycerin. When taken with nitroglycerin, a man's blood pressure can become dangerously low. Some men have died after taking these drugs with nitroglycerin. Drugstore "Even though the study showed that ED does not improve risk prediction beyond the Framingham risk score, an ED assessment can be done at very low cost and presents no risk to patients (unlike other novel CVD screening tests)," said Araujo. "Previous work from our study shows that a simple single question ED measure correlates well with an independent physician's assessment. Therefore, health professionals should consider asking about ED in their older male patients," continued Araujo. "Also, when a patient presents with ED, health professionals should work the patient up for CVD as there may be a window of opportunity for men to improve their health before a CVD event occurs." Although erectile dysfunction is more common in older men, this common problem can occur at any age. "If a man says 'no' to you with a lot of irritability, I still wouldn't take it personally or feel rejected, because it's very likely he's suffering some depression -- and again, it's not you," says Foley. Intraurethral suppositories

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