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Trinucleidae [Trinucleina]
   |--Yinpanolithus Lu in Lu & Chang 1974 [Trinucleinae]
   |    |--*Y. yinpanensis Lu in Lu & Chang 1974
   |    |--Y. guizhouensis Yin in Yin & Li 1978
   |    |--Y. tenuilimbatus (Zhang in Qui et al. 1983)
   |    `--Y. wutangensis (Lu in Wang et al. 1962)
        |--Cryptolithus tesselatus
        |--Salterolithus Bancroft 1929 [incl. Smeathenia Dean 1960]
        |    |--*S. caractaci (Murchison 1839) (see below for synonymy)
        |    `--S. harnagensis (see below for synonymy)
        `--Broeggerolithus Lamont 1935 (see below for synonymy)
             |--*B. broeggeri (Bancroft 1929) [=Cryptolithus broeggeri, *Broeggeria broeggeri]
             |    |--B. b. broeggeri [incl. B. constrictus Bancroft in Dean 1960]
             |    |--B. b. globiceps (Bancroft 1929) (see below for synonymy)
             |    |--B. b. soudleyensis (Bancroft 1929) (see below for synonymy)
             |    `--B. b. ulrichi (Bancroft 1949) [=Salterolithus (Ulricholithus) ulrichi]
             `--+--B. discors (Angelin 1854) [=Trinucleus discors, Cryptolithus dicors (l. c.)]
                |--B. nicholsoni (Reed 1910) (see below for synonymy)
                `--B. transiens (Bancroft 1929) [=Cryptolithus transiens, Broeggeria transiens]

Broeggerolithus Lamont 1935 [=Broeggeria Bancroft 1933 (preoc.); incl. Ulricholithus Bancroft 1933 (n. n.)]

Broeggerolithus broeggeri globiceps (Bancroft 1929) [=Cryptolithus globiceps, B. nicholsoni globiceps]

Broeggerolithus broeggeri soudleyensis (Bancroft 1929) [=Cryptolithus soudleyensis, Broeggerolithus soudleyensis]

Broeggerolithus nicholsoni (Reed 1910) [=Trinucleus nicholsoni; incl. Cryptolithus longiceps Bancroft 1929, Broeggerolithus nicholsoni longiceps, B. melmerbiensis Dean 1962, B. simplex Dean 1960]

Salterolithus caractaci (Murchison 1839) [=Trinucleus caractaci; incl. T. intermedius Wade 1911, Salterolithus intermedius]

Salterolithus harnagensis Bancroft 1929 [=Broeggerolithus harnagensis; incl. Salterolithus caractaci var. paucus Cave 1957, Sa. praecursor Dean 1960, Sa. smeathenensis Bancroft in Dean 1958, Smeathenia smeathenensis]

* Type species of genus indicated

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Bowdler-Hicks, A., J. K. Ingham & A. W. Owen. 2002. The taxonomy and stratigraphical significance of the Anglo-Welsh Cryptolithinae (Trinucleidae, Trilobita). Palaeontology 45 (6): 1075-1106.

Prothero, D. R. 1998. Bringing Fossils to Life: An introduction to paleobiology. WCB McGraw-Hill: Boston.



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