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The Cheirolepidiaceae were a family of conifers, superficially similar to , that were a significant part of the flora (Axsmith et al., 2004). They are united by the possession of a distinctive pollen type assigned to the . The name (as a separate family) or "frenelopsids" has been used for a group of Cheirolepidiaceae with jointed stems, thick , sheathing leaf bases and reduced free leaf tips.

Note that while has been listed below as a synonym of in order to reflect an established association between these taxa, this is not technically correct. Both are "", the former referring to the shoots, the latter to the pollen cones. Because organ genera only refer to specific parts of the plant, they cannot usually be officially synonymised. The use of organ genera has developed in because plant fossils are most often found with the different parts separated from each other. Not only is it not always possible to confidently establish which cone goes with which shoot, but different species may not differ in every part – for instance, different shoots assignable to have been found bearing cones assignable to both and (Axsmith et al., 2004)

Cheirolepidiaceae [Frenelopsidaceae]
   |--Alvinia bohemica
   |--Tomaxellia biforme
   |--Otwayia Pole 2000
   |    |--*O. tetragona (Cantrill & Douglas) Pole 2000 (see below for synonymy)
   |    `--O. cudgeloides Pole 2000
   |--Classostrobus Alvin, Spicer & Watson 1978 [incl. Pseudofrenelopsis, Suturovagina]
   |    |--*C. comptonensis Alvin, Spicer & Watson 1994 (see below for synonymy)
   |    |--C. arkansensis Axsmith, Krings & Waselkov 2004
   |    |--C. cathayanus [incl. Pseudofrenelopsis intermedia]
   |    |--C. turolensis Gomez, Martín-Closas et al. 2002
   |    `--C. ugnaensis Gomez, Martín-Closas et al. 2002
   |--Classopollis Pflug 1953
   |    |--*C. classoides Pflug 1953
   |    |--C. brasiliensis Herngreen 1975
   |    |--C. mirabilis Reyre 1970
   |    |--C. noeli Reyre 1970
   |    `--C. obidosensis Groot & Groot 1962
   |--Cupressinocladus Seward 1919 [incl. Watsoniocladus]
   |    |--C. asferensis Barale & Ouaja 2002
   |    |--C. itieri (Saporta) Barale 1981
   |    |--C. leptocladoides (Berry) Pons 1990
   |    |--C. micromerum (Heer) Pais 1974
   |    |--C. pompeckji (Salfeld) Pons 1990
   |    |--C. pseudoexpansum Barnard & Miller 1976
   |    `--C. soltanensis Barale, Ouaja & Philippe 2000
   `--Frenelopsis Schenck 1869
        |--*F. hoheneggeri (Ettingshausen) Schenck 1869 [=Thuites hoheneggeri Ettinghausen 1852]
        |--F. alata Feistmantel 1881
        |--F. choshiensis
        |--F. elegans
        |--F. harrisii Doludenko & Reymanówna 1978
        |--F. kaneviensis Barale & Doludenko 1985
        |--F. occidentalis
        |--F. oligostomata [incl. Classoidites glandis]
        |--F. pombetsuensis Saiki 1997
        |--F. ramosissima
        |--F. rubiesensis Barale 1973
        |--F. sifloana Watson 1983
        |--F. teixeirae
        |--F. turolensis Gomez, Martín-Closas et al. 2002
        |--F. ugnaensis Gomez, Martín-Closas et al. 2002
        `--F. veneta Gomez, Thévenard et al. 2002

*Classostrobus comptonensis Alvin, Spicer & Watson 1994 [incl. Classopollis martinottii, Pseudofrenelopsis parceramosa (Fontaine) Watson 1977 (mt)]

*Otwayia tetragona (Cantrill & Douglas) Pole 2000 [=Geinitzea tetragona Cantrill & Douglas 1988]

* Type species of generic name indicated

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Axsmith, B. J., M. Krings & K. Waselkov. 2004. Conifer pollen cones from the Cretaceous of Arkansas: Implications for diversity and reproduction in the Cheirolepidiaceae. Journal of Paleontology 78: 402-409.

Barale, G., & M. Ouaja. 2002. La biodiversité végétale des gisements d’âge Jurassique supérieur-Crétacé inférieur de Merbah El Asfer (Sud-Tunisien). Cretaceous Research 23: 707-737.

Gomez, B., C. Martín-Closas, G. Barale, N. Solé de Porta, F. Thévenard & G. Guignard. 2002. Frenelopsis (Coniferales: Cheirolepidiaceae) and related male organ genera from the Lower Cretaceous of Spain. Palaeontology 45 (5): 997-1036.

Gomez, B., F. Thévenard, M. Fantin & L. Guisberti. 2002. Late Cretaceous plants from the Bonarelli Level of the Venetian Alps, northeastern Italy. Cretaceous Research 23: 671-685.

Ibrahim, M. I. A. 2002. Late Albian-Middle Cenomanian palynofacies and palynostratigraphy, Abu Gharadig-5 well, Western Desert, Egypt. Cretaceous Research 23: 775-788.

Pole, M. S. 2000. Mid-Cretaceous conifers from the Eromanga Basin, Australia. Australian Systematic Botany 13 (2): 153-197.


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