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Taxonomy Phylogeny

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Crocodylomorpha


The diminutive crocodylomorph, (,,), of late Triassic () Wales, pursuing a . (c) Stanton F. Fink)


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The following section of this page needs to be . At its simplest, wikifying can just mean adding links to other pages via the edit tab. For help in editing Palaeos org pages, see the page

Today's crocodiles are mostly large reptiles are adapted to a semi-aquatic existence. This was not always the case. In fact the crodylomorph group began as small, gracile terrestrial predators. Crocodile scales are hard and square, and a few rows of raised scales down the back and tail contain knobs of bone. Every bit as old as the dinosaurs, to which they are fairly closely related, the crocodiles and their early predecessors (collectively, "Crocodylomorpha") are an ancient and venerable lineage of archosaurs. Although typically reptilian in their scaly integument and "cold blooded" (ectothermic - actually inertial homeothermic in large forms) metabolism, they nevertheless display advanced chracteristics such as a four-chambered heart (in efficency similiar to the bird and mammal heart) and maternal care of young.

During the late Triassic and early Jurassic these animals were mostly small, graceful, active terrestrial forms, quite unlike the way we think of crocodiles today. After flourishing for some 40 or 50 million years - almost as long as the age of mammals - these small active animals died out, either through competition with small theropod dinosaurs, or perhaps through a mass extinction (e.g. the that also eliminated the Prosauropod, Coelophysoid and Heterodontosaur dinosaurs and a number of lines of marine reptiles and invertebrates). The surviving crocodilians were larger animals, more like the crocs we know today, but, curiously, mostly marine. Later in the Jurassic, these marine forms invaded fresh water and swamp environments as large semi-aquatic predators, and finally returned to the land again as small forms.

It was as large, semi-aquatic predators that they became most successful. Crocodiles are superbly adapted to life in rivers and swamps. A fold of skin closes the windpipe at the back, so that the animal can open its mouth under water and breathe with only its nostrils above the surface. The nostrils, like the eyes and ears, are placed high on the head. When crocodiles are completely submerged, the ears and nostrils are closed by valves, and the eyes covered by membranes. Crocodiles have no salivary glands and usually eat under water. Some species sweep their prey from the land into the water with their powerful, vertically flattened tails, a technique that was almost certainly also practiced by Phytosaurs of the late Triassic.

The warm balmy Mesozoic climate was kind to these prehistoric reptiles, and they flourished and diversified for many millions of years, attaining world-wide distribution. The larger semi-aquatic types lunched on dinosaurs and large lungfish and Lepidotids. Some, like the great Sarcosuchus and Phobosuchus, reached huge size, 10 to 15 meters in length and some 8 to 15 tonnes in weight, bigger than the largest T. rex.

Perhaps because of their ectothermic metabolism (which makes it possible to go without food for long periods) and their reliance on the fresh water carrion - detrivore foodchain, rather than the ocean plankton - based or the terrestrial plant - based foodchains, the crocodiles were unaffected by the comet/asteroid - impact end-Cretaceous mass extinction that eliminated most forms of life on Earth. They continued through to the Paleocene, and in the absence of large competitors evolved into large fully terrestrial preditors (Sebecosuchids, Pristochampsids).

The global cooling trend that began in the Oligocene epoch and accelerated in the Miocene and Pliocene was catastrophic for the crocodiles. Many types died out, unable to cope with the increasing cold, and the survivors retreated to the tropics and a life in swamps. The start of the modern epoch (Holocene epoch) saw 21 species distributed in (depending on how we draw the lines) one, two, or three families (four subfamilies).

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(The following menu and phylogeny refers to subtopics of this page)


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Taxonomy Phylogeny

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Phylogeny:  ::: + Crocodylomorpha : + .

:

  • Phylum:
    • Class: (traditional) /
      • Infraclass:
        • Superorder: Crocodylomorpha

Author: , 1930

Stratigraphic Range: Middle Triassic () to .

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The Crocodylomorpha are defined by Sereno 2005 as "The most inclusive containing (Laurenti 1768) but not Mehl 1915, Romer 1972, Huene 1942, Fraas 1877."

This a definition and therefore includes all taxa closer to extant crocodilians than to other clades.

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The following classification is from the Bristol University webpage on Crocodylomorpha. All these taxa are extinct except for the three Eusuchian shown here. There are in addition many other, less well-known groups that are not included in this current listing. provides a more detailed

  • Division
    • Superorder Crocodylomorpha
      • Family
      • Family
      • Order ()
        • Family
        • Division
          • Family
          • Family
          • Subdivision
            • Family
            • Family
            • Infradivision
              • Family
              • Family
              • Suborder
                • Family
                • Family
                • Family

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  • - webpages by Ross Elgin on the server
    • classification (used here)
  • - hyperlinked cladogram at

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The following section of this page needs to be . At its simplest, wikifying can just mean adding links to other pages via the edit tab. For help in editing Palaeos org pages, see the page

Characters: Skull massive and well-buttressed; no skull kinesis (skull absorbs force of strong, rapid bite); antorbital fenestrae small (closed in modern forms); descending process of prefrontal present [π$03]; postfrontals absent; descending process of squamosal absent [$P93] [π$03]; squamosal overhangs temporal region [π$03]; anterior extension of quadrate; quadrate complex; quadrate contacts prootic [π$03]; occiput solid, no or minor occipital fossae [π$03]; exoccipital with lateral crest lying anterior to hypoglossal foramen [$G02]; palate & braincase strongly integrated; ventral ramus of opisthotic makes extensive contact with prootic anteroventral to fenestra ovalis [π$03]; middle ear elements pneumatized [π$03]; crista vestibuli present [π$03]; lagenar recess visible externally as prominence on wall of otic capsule [π$03]; Eustachian tubes at least partially encased in bone [π$03]; prootic not in broad contact with anterior of paroccipital process [π$03]; external foramen for abducens nerve solely within basisphenoid [π$03]; epipterygoid absent; basipterygoid processes enlarged [π$03]; pila antotica ossified by basisphenoid and laterosphenoid so that these elements are in contact anterior to trigeminal foramen [π$03]; secondary palate formed by maxillae & vomers [$P93]; large anterior & posterior pterygoideus muscles, oriented horizontally; parasphenoid rostrum forms vertically expanded wedge [π$03]; skull bones pneumatized; 24 presacral vertebrae, 2 sacral vertebrae; accessory neural spine on mid-caudal vertebrae [π$03]; cartilaginous sternum (modern forms) separates posteroventral extensions of coracoids; no clavicle (all suggest obligate quadruped); elongated carpals (ulnare and radiale) perhaps involved in giving extra swing to foreleg [$P93]; manus IV with 5 phalanges (reversal) [π$03]; pubic shaft length >3x width of acetabulum [π$03]; pubis excluded from acetabulum since Late Jurassic; acetabulum semi-perforate [π$03]; femoral head without distinct neck, but bent medially with parallel medial & lateral sides [$P93]; metatarsal I length at least 85% length of Mt III [π$03]; pes IV with <5 phalanges [$P93]; pes V with 2 phalanges [π$03]; paramedial osteoderms sculptured [$P93]; osteoderms with anterolateral articulatory process & dorsolateral keel (as in Euscolosuchus) [S92].

Notes: [1] "[π03]" refers to our own analysis of published data.

References: Gower (2002) [G02], Parrish (1993) [P93], Sues (1992) [S92].

ATW031223.

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  • (2004), , 3rd ed.
  • Hay, O. P. 1930 (1929-1930). Second Bibliography and Catalogue of the Fossil Vertebrata of North America. Carnegie Institution Publications, Washington, 1,990 pp.
  • . 2005. [version 1.0, 2005 November 7]

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