Goniatitida

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(provided links back to Ammonoidea and Cephalopoda in text.)
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The Goniatitida constitute an order of [[Ammonoidea|ammonoid]] [[Cephalopoda|cephalopods]] from the late Paleozoic and are found in Middle Devonian to Upper Permian marine sedimnets. The Goniatitida diversified into two main lineages in the Early Carboniferous (Mississippian) and subsequently into a number of sublineages, forming different families and superfamilies.  
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'''Goniatitida''' constitute an order of [[Ammonoidea|ammonoid]] [[Cephalopoda|cephalopods]] from the Late [[Paleozoic]] and are found in Middle [[Devonian]] to Upper [[Permian]] marine sediments. The Goniatitida diversified into two main lineages in the Early [[Carboniferous]] ([[Mississippian]]) and subsequently into a number of sublineages, forming different families and superfamilies.  
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Goniatitids (goniatites) are united by a basic suture of 8 simple lobes, less in primitive forms, more that may be ammonitic in advanced forms, and a [[siphuncle]] that is prosiphonate. (MF&S)  
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Goniatitids (goniatites) are united by a basic suture of 8 simple lobes, less in primitive forms, more in advanced forms that may be ammonitic, and a [[siphuncle]] that is prochoanitic (prosiphonate (MF&S)  
==Morphology==
==Morphology==
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The shells of gonitites are planospirally coiled and vary from thinly discoidal to broadly globular. They may be smooth or distinctly ornamented. Coiling may be evoluate with all whorls exposed, or involute with only the last whorl showing.  
The shells of gonitites are planospirally coiled and vary from thinly discoidal to broadly globular. They may be smooth or distinctly ornamented. Coiling may be evoluate with all whorls exposed, or involute with only the last whorl showing.  
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==Classification.==  
==Classification.==  
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The Goniatitida are divided into two suborders, the [[Tornoceratina]] and the [[Goniatitina]] which represent the two main evolutionary groups within the order shown in the Treatise (Part L). GONIAT online, shows six superfamilies in the Tornoceratina and 16 within the Goniatitina.  
The Goniatitida are divided into two suborders, the [[Tornoceratina]] and the [[Goniatitina]] which represent the two main evolutionary groups within the order shown in the Treatise (Part L). GONIAT online, shows six superfamilies in the Tornoceratina and 16 within the Goniatitina.  
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===Nomenclatural note===
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===<small>Phylogeny in the Treatise-1957</small>===
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Note that in the GONIAT website, superfamilies end in --oidea, conforming wth the recent ICZN ruling on superfamily names necessitating taxonomic rank be specified. Previously in invertebrate paleontology names ending in --oidea were limited (mostly) to subclasses and superorders and --aceae was universally used for superfamilies, eleminating the need to specify rank. Most recent taxonomies and revisions have adopted the ICZN ruling (recommondation?)
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The Treatise Part L, 1957, indicates in fig. 12, Phylogeny of Paleozoic Ammonoids, two primary lineages stemming from the Tornoceratidae; 1. the Tornoceratid, beginning with the Tornoceratidae and 2. the Goniatitid, beginning with the Goniatitidae. The Tornoceratidae, accordingly, gave rise to the Cheiloceratidae from which the  Agathiceratidae, followed by the Shumartitidae, Popanoceratidae, Perrinideae and Cyclolobiidae are subsequently derived.  The Goniatitidae, which seems to have its origin the the Cheiloceratidae  gave rise to the Neoicoceratidae and Schistoceratidae which are followed by the Metalegoceratidae and Adrianitidae, and also gave rise to the Dimorphoceratidae which in turn gave rise to the Thalassoceratidae.
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==''References''==
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The Tornoceratidae and Cheiloceratidae are combined in the the Cheilocerataceae, the Agathiceratidae, Shumarditidae, and Perrinitidae in the Agathicerataceae; and the Popanoceratidae and Cylolobiidae in the Cyclolobaceae. The three superfamilies were the original Tornoceratina. 
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The Goniatitidae, with subfamilies, Neoicoceratidae, Metalegoceratidae and Schistoceratidae are combined in the Goniatitaceae; the Adrianitidae as its own superfamily, Adrianitaceae; the Dimophoceratidae and Thalassoceratidae in the Dimorphocerataceae.  These three superfamilies were the original Goniatitina.
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===<small>Subsequent developments -Current persepctives</small>===
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One of the more significant developments since the first edition of the Treatise, part L, is the elevation of the Goniatitina to full ordinal status as the Goniatitida, which allowes phylogenetically related superfamilies to be brought together as the  suborders Tornoceratina  and Goniatitna.
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Another is the use of the --oidea ending for superfamlies by some, as proscribed by the ICZN. More significant however are the rearrangements and addition of new families and superfamilies.
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==''References''==
*Miller, Furnish, and Schindewolf (M F &S); Paleozoic Ammonoidea, in The Treatise on Invertebrate Paleontology, Part L, Ammonoidea; Geological Society of America and University of Kansas press, 1957.
*Miller, Furnish, and Schindewolf (M F &S); Paleozoic Ammonoidea, in The Treatise on Invertebrate Paleontology, Part L, Ammonoidea; Geological Society of America and University of Kansas press, 1957.
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*[http://www.goniat.org/index.html|GONIAT]GONIAT Online.
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*[http://www.goniat.org/index.html GONIAT Online]
[[user:John M|JHM]] 8/17/09
[[user:John M|JHM]] 8/17/09

Latest revision as of 17:32, 22 June 2011

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