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Jump to: ,
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Halkieria
from the , North Greenland

Halkieria is a fossil organism from the to . It has been found on almost every continent in deposits of this age in socalled assemblages.

Halkieria is best known from the North Greenland in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal later a more thorough description was undertaken in 1995 in the journal and larger evolutionary implications was posed.

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The animal was bilaterally symmetric, dorso-ventrally flattened with a naked ventral side. The dorsal side was covered with imbricating in three zones. In either end, there is a larger shellplate with prominent growthlines. The sclerites are distinguished into three types; the palmate sclerites placed between the two shellplates, surrounding these there is a zone of sclerites termed the cultrates and along the entire margin, there is a zone of spineshaped sclerites called siculates. The shellplates and the sclerites was originally of calcium carbonate.

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When Conway Morris and Peel described from North Greenland they noticed the similarity to known especially from the which has sclerites organised in similar zones. They also noticed the similarity of the shellplates to inarticulate . This led them to suggest that Halkieria is a stemgroup brachiopod which had evolved into the bivalved brachiopod by gradually getting inclosed within the two valves and starting a lifemode. Wiwaxia would be the stemgroup of or according to Nicholas Butterfield (1990) which is led by the recognition of its sclerites having a similar ultrastructure as polychaete chaetae secreted by numerous microvillae.

A number of other scientists has been stressing affinites of Halkieria. According to Vinther and Nielsen (2005) there is no characters supporting either brachiopod or annelid affinity of Halkieria. All characters seem to be compatible with molluscs, especially the chitons or .

Lately, a redescription of the fossil has revealed it as being a mollusc relative instead of some sort of . The tooth apparatus has many compelling similarities with a molluscan along with a foot-like area and surrounding gill-like structures (supposed ). The tooth apparatus is also very similar to , which hereby possibly marks some important links between these .

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  • Vinther, J. and Nielsen, C. (2005) The Early Cambrian Halkieria is a mollusc, Zoologica Scripta, 34 (1) 81-89

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