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The Late Jurassic Epoch of the Jurassic Period
161 to 146 million years ago

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The late Jurassic period sees the evolution of some of the greatest of all. The continue to flourish and to diversify, as the older Mid-Jurassic are replaced by a diverse Late Jurassic fauna of , , and . Some of these creatures attained tremendous size. The giraffe-like Brachiosaurus reached 22 metres and weighed 15 tonnes and more. Seismosaurus hallorum (AKA Diplodocus hallorum), a more slender animal, exceeded 40 metres in length.

In addition there were many other types of animals around at this time: a number of different types of , , , , , flying reptiles (), marine reptiles, and the first ().

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EpochAgeBase (Mya)Duration (Ma)
 Late Jurassic 
 (Malm Epoch) 

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Much can be gleaned of the biogeography of this ancient world by studying the distribution of the dinosaurs. Four localities of importance as regards fossil remains are known; central North America (the of the South-West), East Africa ( of Mkwoa wa Mtwara, Tanzania), Europe (scattered localities mainly in England, France, Bavaria, and Portugal), and China (especially the of Sichuan Province). Smaller vertebrates and invertebrates are also known in abundance from France and Bavaria. A comparison of the types of creatures from these localities shows often the same family, and sometimes even the same genus, is represented in different localities, showing that, just as in the preceding period, the world was a much more biotically uniform place than it is now. This is due to the fact that all the continents had merged into a single landmass, . Yet high sea levels broke this supercontinent into a series of large islands, causing regional differences and faunal as well as floral provinces.

The following faunal (and megafaunal) provinces can be tentatively distinguished:

West Laurasia: Probably everyone is familiar with the great American dinosaurs of the Jurassic: the huge Apatosaurus (AKA "Brontosaurus"), its slender cousin Diplodocus, the large carnivores Allosaurus and Ceratosaurus, and the striking plated Stegosaurus. Alongside these were a number of smaller herbivores and carnivores (Camptosaurus, Dryosaurus, Ornitholestes, etc). In this megafauna sauropods predominated, and were primarily of the diplodocid and camarasaurid families, with a few haplocanthosaurs (especially early on) and the occasional huge brachiosaur (in fact the brachiosaurs were bigger here than anywhere else). As Dr Bob Bakker has shown, the West Laurasian dinosaurs exhibited a tendency to increase in size, possibly caused by increasingly harsh and arid conditions, so that by the mid-Tithonian they were truly gigantic. Then they suddenly died out, leaving a number of small dinosaurs, mammals, and lower vertebrates to carry on.

West Gondwanaland: West Gondwanaland (e.g. Africa) had a fauna very similar to that of North America, with many types of dinosaurs in common. The theropod Ceratosaurus, the diplodocid Barosaurus, and the ornithopod Dryosaurus occur on both continents. But the Laurasian Camarasaurus is absent, and the mighty brachiosaurs, which is rare in North America (West Laurasia), is the most common sauropod in the Tendaguru formation (West Gondwana). However, it appears to be a distinct genus (designated Giraffatitan) from the West Laurasian form. The small stegosaur Kentrosaurus closely resembles primitive European and Chinese forms, but is quite unlike the large and advanced American Stegosaurus. Pterosaurs are known too, these belong to the same genus (Rhamphorhynchus) that inhabited the islands of Europe. There were also several types of dinosaurs that were Gondwanan endemics as well. Of significance were Dicraeosaurus (a small short-necked persistently primitive diplodocoid sauropod), Torniera (a medium-sized sauropod, possibly ancestral to the Cretaceous ), and the poorly known ceratosaur Elaphrosaurus. These three taxa indicate the development of a unique Gondwanaland fauna.

Central Laurasia: Whereas Africa and North America were large landmasses, Europe during the Jurassic seems to have consisted of a number of large islands separated by shallow sea. A number of different types of dinosaurs have been found in Portugal. They include not only most "standard" Jurassic types (brachiosaurids, stegosaurs, megalosaurs, etc) but also small (armoured dinosaurs)). As with the early Jurassic , this latter group are almost totally indigenous to Europe (which may well be the centre of evolution of these armoured dinosaurs) during the Jurassic period. But during the following early Cretaceous period they spread throughout the globe. The first protobirds like Archaeopteryx also inhabited the islands of Europe, no doubt benefiting from the isolation from large predators and tendency to evolutionary novelty that small islands provide.

East Laurasia and East Gondwana: Animals on the eastern part of the world-continent remained quite different to those of the west. Stegosaurs were exceedingly diverse in China, and include both primitive and advanced forms. They lived alongside basal (ancestral) , persistently primitive euhelopid sauropods and Triassic holdovers like tritylodontid and short-headed brachyopid . Similar chigutosaurid temnospondyls must have lived in East Gondwanaland (Australia) as well, because they are known there from both the Early Jurassic and the middle Cretaceous. Allosaurids are also known from the Late Jurassic of China and the middle Cretaceous of Australia, indicating that these carnivores were able to wander widely over the Earth's surface (unlike the euhelopid sauropods which are not known outside of China).


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