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Plankton


Ceratium, a phytoplankter


Plankton are drifting organisms that inhabit the of , , and bodies of .

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Some marine - a key group

The name plankton is derived from the word πλανκτος ("planktos"), meaning "wanderer" or "drifter" (Thurman, 1997). While some forms of plankton are capable of independent movement and can swim up to several hundreds of metres vertically in a single day (a behavior called ), their horizontal position is primarily determined by in the body of water they inhabit. By definition, organisms classified as "plankton" are unable to resist ocean currents. This is in contrast to , which are organisms that can swim against the ambient flow of the water environment and control their position (e.g. , , and ).

Within the plankton itself, are those organisms that spend their entire as part of the plankton (e.g. most , , , and ). By contrast, are those organisms that are only planktonic for part of their lives (usually the larval stage), and then graduate to either the nekton or a existence. Examples of meroplankton include the larvae of , , , marine , and most .

Plankton abundance and distribution are strongly dependent on factors such as ambient concentrations, the physical state of the water column, and the abundance of other plankton.

The study of plankton is termed planktology. Individual plankton are referred to as plankters.

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An (Hyperia macrocephala)

Plankton are primarily divided into broad functional (or ) groups:

  • (from Greek phyton, or plant), or that live near the water surface where there is sufficient to support . Among the more important groups are the , and .
  • Zooplankton (from Greek zoon, or animal), small or (e.g. and other ) that feed on other plankton. Some of the and of larger animals, such as fish, crustaceans, and , are included here.
  • , and , which play an important role in organic material down the water column (note that many phytoplankton are also bacterioplankton).

This scheme divides the plankton community into broad producer, consumer and recycler groups. In reality, even the trophic level of some plankton is not straightforward. For example, although most dinoflagellates are either photosynthetic producers or heterotrophic consumers, many species are depending upon their circumstances.

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Plankton are also often described in terms of size. Usually the following divisions are used:

  • Megaplankton, 2×10-1→2×100 m (20-200 cm)
  • Macroplankton, 2×10-2→2×10-1 m (2-20 cm)
  • Mesoplankton, 2×10-4→2×10-2 m (0.2mm-2 cm)
  • Microplankton, 2×10-5→2×10-4 m (20-200 µm)
  • Nanoplankton, 2×10-6→2×10-5 m (2-20 µm)
  • Picoplankton, 2×10-7→2×10-6 m (0.2-2 µm), mostly
  • Femtoplankton, < 2×10-7 m, (< 0.2 µm), consisting of marine

However, some of these terms may be used with very different boundaries, especially on the larger end of the scale. The existence and importance of nano- and even smaller plankton was only discovered during the , but they are thought to make up the largest proportion of all plankton in number and diversity.

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, a
Plankton are found throughout the oceans, seas and lakes of Earth. However, the local abundance of plankton varies horizontally, vertically and seasonally. The primary source of this variability is the availability of light. All plankton ecosystems are driven by the input of solar energy (but see ), and this confines primary production to surface waters, and to geographical regions and seasons when light is abundant.

A secondary source of variability is that of nutrient availability. Although large areas of the and oceans have abundant light, they experience relatively low primary production because of the poor availability of nutrients such as , and . This is a product of large-scale and of the water column. In such regions, primary production, still usually occurs at greater depth, although at a reduced level (because of reduced light).

While plankton are found in the greatest abundance in surface waters, they occur throughout the water column. At depths where no primary production occurs, zooplankton and bacterioplankton instead make use of organic material sinking from the more productive surface waters above. This flux of sinking material can be especially high following the termination of spring blooms.

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Plankton play a role in the of many important . Of particular contemporary significance is their role in the ocean's .

As stated, phytoplankton fix in sunlit surface waters via photosynthesis. Through (primarily) zooplankton grazing, this carbon enters the planktonic foodweb, where it is either to provide energy, or accumulates as or . As living or dead organic material is typically more than seawater it tends to sink, and in open ocean ecosystems away from the coasts this leads to the transport of carbon from surface waters to the deep. This process is known as the , and is one of the reasons that the oceans constitute the largest (active) pool of carbon on .

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  • Omori, M., T. Ikeda (1992). Methods in Marine Zooplankton Ecology. Malabar, USA: Krieger Publishing Company.
  • Thurman, H. V. (1997). Introductory Oceanography. New Jersey, USA: Prentice Hall College.

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