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Revision as of 09:54, 19 October 2008 by ( | )
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Pteraspidomorphi

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Pteraspidomorphi is an extinct of . The fossils show extensive shielding of the head. Maybe some species even lived in . They are considered to be the most primitive of vertebrates.

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The Pteraspidomorphi are sometimes referred to as Heterostraci, although that term is more properly applied to the specialised and forms. They are an extinct of jawless fish, and were the most abundant and diverse vertebrates of the Silurian. They are distinguished by having an or bony shield composed of several plates, usually a (upper or back) shield, a (lower or belly) shield, branchial (gill) plates, and a number of smaller plates around the areas of the mouth (oral plates) and eyes. These plates consist of a mostly acellular form of bone, called , which is believed to be the ancestral condition for the dermal or skin/exoskeleton bone of all bony vertebrates. The body is covered by many scales, each of which has ornament matching the type seen on the larger dermal bones. In all of the more derived forms, there is only one, common gill opening on each side.

The Pteraspidomorphi include the earliest known vertebrates. The oldest certain remains date to the or , although the disputed and fragmentary Anatolepis probably goes back to the late Cambrian. In any case, by the middle and late Ordovician, there were several different lineages evolving in isolation different parts of the world, as this map indicates.

These Ordovician forms, the Arandaspida, Astraspida and Eriptychiida, were formerly grouped with the Silurio-Devonian Heterostraci, but are rather more primitive. These early types share some unique features with heterostracans, such as the presence of large median dorsal and ventral plates, but do not possess common external branchial openings.

By the start of the Silurian period these lineages had died out, possibly as another result of the mass extinction at the end of the Ordovician. They were replaced by the Heterostraci. This latter group underwent an , dividing into a number of lineages and reaching their peak during Late Silurian and Early Devonian times, when a variety of different types evolved and flourished, from to . All had the characteristic head shield, which could grow throughout the life of the animal.

By the middle Devonian the Pteraspidomorphi went into decline, with only a single family of giant (by agnath standards) flattened bottom-dwellers, the psammosteids, continuing almost until the end of the Devonian, the last and also the largest of the armoured jawless fish.

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Loosely synonymous with "Diplorhina". Probably defined as Anatolepis + Psammolepis.

Range: (or ) to .

Phylogeny:  ::: "" + Pteraspidomorphi : + ( + ).

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: 10-200 cm; head very long (40-50% body length); paired nasal capsules, and ?openings; solid anterior plates, with at least a major oblong median dorsal and ventral plates and usually a median rostral plate above mouth; rostra and cornua common; pineal opening (psammosteids?); brain with two semicircular canals; some show internal impressions of arcualia on notochord; body tends to be ventrally flattened (numerous exceptions); mobile tail, generally fan- or pad-like; tail hypocercal; no paired appendages; no unpaired fins except caudal; sensory line system present; no cellular bone (but see [H73]); possible traces of calcified cartilage in in some basal lineages; trilaminate aspidine; grebeshki ornamentation on dermal plates (lost independently several times); largely marine nearshore.

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o †Pteraspidomorphi Goodrich, 1909
  |-- †
  `--+--o † Ritchie & Gilbert-Tomlinson, 1977
     |  |?- † rinconensis Erdtmann et al., 2000
     |  |--o † Gagnier, Blieck & Rodrico, 1986 [Sacabambaspidae]
     |  |  |-- †S. sp. [Young, 1997] [Ordovician, Central Australia]
     |  |  `-- †S. janvieri Gagnier, Blieck & Rodrico, 1986
     |  `--o † Ritchie & Gilbert-Tomlinson, 1977
     |     |--o † Ritchie & Gilbert-Tomlinson, 1977
     |     |  |-- †A. sp. [Young, 1997] [Ordovician, Central Australia]
     |     |  `-- †A. prionotolepis Ritchie & Gilbert-Tomlinson, 1977
     |     `--o † Ritchie & Gilbert-Tomlinson, 1977
     |        |-- †P. sp. [Young, 1997] [Ordovician, Central Australia]
     |        `-- †P. crenulata Ritchie & Gilbert-Tomlinson, 1977
     `--o † Lankester, 1868
        |-- † desiderata Walcott, 1892 [taxonomic history] [Astraspida Berg, 1940: Astraspidiformes Berg, 1940: Astraspidae Eastman, 1917]
        `--+-- † americanus Walcott, 1892 [taxonomic history] [Eriptychiida: Eriptychiidae Tarlo, 1962:]
           `--o † Lankester, 1868
              |?-o †
              |  `-- †A. canadensis Dineley & Loeffler, 1976
              |--o †
              |  `-- †L. serrata Dineley & Loeffler, 1976
              |?- †
              |?- †
              |?- †
              |?- †
              `--+--o †Traquairaspidiformes
                 |  `--o †Traquairaspididae
                 |     |--o †
                 |     |  `-- †P. symondsi
                 |     |-- † [Lophopiscis, Lophaspis, Orthaspis, Phialaspis, Yukonaspis]
                 |     `--o †
                 |        |-- †W. godmani Tarlo
                 |        `-- †W. brotzeni Tarlo, 1964
                 `--+--o †Tolypelepidida Denison, 1964 [Cyathaspididae: Tolypelepidinae Denison, 1964]
                    |  `--o †Tolypelepididae Denison, 1964 [Tolypelepidinae Denison, 1964]
                    |     |-- † interstincta Dineley & Loeffler, 1976
                    |     |-- † chattertoni Soehn & Wilson, 1990
                    |     `--o † Pander, 1856 [Tolypaspis]
                    |        |-- †T. undulata Pander, 1856
                    |        |-- †T. lenzi Dineley & Loeffler, 1976
                    |        `-- †T. leopoldensis Loeffler & Jones, 1977
                    `--+-- †
                       `-- † Berg, 1940

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  • Halstead, LB (1973), The heterostracan fishes. Biol. Rev. 48: 279-332. [H73]

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